PTERIDOPHYTES OF UPPER KATANGA (DEMOCRATIC REPUBLIC OF CONGO) PAPROTNIKI GÓRNEJ KATANGI (REPUBLIKA DEMOKRATYCZNA KONGA) KRAKÓW 2000 BOTANICAL PAPERS 35 JAN KORNAS ANNA MEDWECKA-KORNAŚ FRANÇOIS MALAISSE MAŁGORZATA MATYJASZKIEWICZ PTERIDOPHYTES OF UPPER KATANGA (DEMOCRATIC REPUBLIC OF CONGO) PAPROTNIKI GÓRNEJ KATANGI (REPUBLIKA DEMOKRATYCZNA KONGA) KRAKÓW 2000 THE INSTITUTE OF BOTANY OF THE JAGIELLONIAN UNIVERSITY PRACE BOTANICZNE 35 JAN KORNAŚ ANNA MEDWECKA-KORNAŚ FRANÇOIS MALAISSE MAŁGORZATA MATYJASZKIEWICZ PTERIDOPHYTES OF UPPER KATANGA (DEMOCRATIC REPUBLIC OF CONGO) PAPROTNIKI GÓRNEJ KATANGI (REPUBLIKA DEMOKRATYCZNA KONGA) KRAKÓW 2000 NAKŁADEM INSTYTUTU BOTANIKI UNIWERSYTETU JAGIELLOŃSKIEGO Redaktor Anna Stengl Editor Sekretarz Redakcji Małgorzata Matyjaszkiewicz Editorial Assistant Recenzenci * Reviewers Ryszard Ochyra Rada Redakcyjna • Editorial Council Zbigniew Dzwonko, Małgorzata Matyjaszkiewicz, Maria Olech, Anna Stengl, Adam Zajac (przewodniczący - president), Alicja Zemanek, Bogdan Zemanek Skład komputerowy Projekt okładki Marian Wysocki Maciej Pierzchała Make-up Editor Cover Design Adres Redakcji • Editorial Instytut Botaniki Uniwersytetu Jagiellońskiego ul. Lubicz 46, 31-512 Kraków Institute of Botany of the Jagiellonian University Lubicz 46, 31-512 Kraków copyright © Instytut Botaniki Institute of Botany Uniwersytetu Jagiellońskiego of the Jagiellonian University Printed in Poland ISBN 83-909074-8-8 ISSN 1509-9156 Zakład Poligraficzny Uniwersytetu Jagiellońskiego Kraków, ul. M. Kopernika 31 Jan Kornas 1923-1994 collecting a hanging club-moss Lycopodium dacrydioides Bak. in Uluguru Mountains - Tanzania, phot. T. Poes Contents Abstract........................................................... 7 Introduction....................................................... 9 1. The area studied and main types of vegetation ................. 9 1. 1. Location and oro-hydrography 9 1. 2. Climate................................................. 12 1. 3. Geology................................................. 12 1. 4. Soils................................................... 14 1. 5. Vegetation.............................................. 15 2. Methods, material and scope of information.................... 20 3. List of species............................................... 24 4. Final remarks on the pteridophyte flora of Upper Katanga.... 130 4. 1. Taxonomic composition ................................. 130 4. 2. Comparison with other areas and some conclusions....... 131 5. Distributional maps.......................................... 135 References ...................................................... 171 Streszczenie..................................................... 175 Appendix......................................................... 178 Index to generic names 181 Abstract - The collection, essential for the present study, was mostly made by Anne Bodenghien and François Malaisse between 1985 and 1988 in Upper Katanga, south-eastern part of the Democratic Republic of Congo, previously the Upper Shaba in Zaire. Voucher specimens sent to the Jagiellonian University Herbarium (KRA) in Kraków, Poland, were worked out by Jan Komaś who, unfortunatelly, passed away on 8 August 1994. In the present publication, prepared thereafter mainly by A. Medwecka-Komaś, the material from KRA and data existing in cited literature were taken into account. In the description of the study area (Figs 1-4) particular attention is paid to vegetation. Some methodological remarks, the comparison of fem family classification by various authors (Table 1) and abbreviations used in the text are given. The floristic list contains information about the distribution and habitat of particular taxa. Their stations are ordered and mapped using the grid method (Figs 5-202). Habitat descriptions are established according to the herbarium labels. More detailed information are to be found in particular citations appearing in the publication concerning fem and fern allies in Zambia (Kornaś 1979). In the final chapter some numerical features of the pteridophyte flora of Upper Katanga are discussed. The list published by the present authors numbers 183 species, including 6 being endemic to the Zambezi - Congo watershed and one, Actiniopteris kornasii, described as new by Medwecka-Kornaś (1999). For the general idea regarding the richness of fern and fern allies in Upper Katanga some comparisons with other regions are presented (Table 2). The flora under consideration comprises a higher number of species than e.g. the pteridophyte flora of larger Zambia, with which 125 species are in common, and similar number of species like very small Rwanda, which has however very different geographical conditions. Data in the present study are adequate for further interpretations, e.g. about the patterns of local species distribution in Upper Katanga and about the presence of various geobotanical elements. Introduction The present publication provides information about the occurrence of fem and fern allies in Upper Katanga, which forms the south-eastern part of the Democratic Republic of Congo (in recent decades, until 1997, called Upper or Meridional Shaba in Zaire). The publication was prepared with the cooperation of Polish and Belgian botanists, in part during two visits of A. Medwecka--Komaś in Brussels. It is based chiefly on the collection of Anne Bodenghien and François Malaisse made in Upper Katanga mostly between 1985 and 1988 and worked out in the Institute of Botany of the Jagiellonian University in Kraków (Poland) by Jan Kornaś, author of numerous publications on African ferns and fern allies. Unfortunately Jan Kornaś, meritorious and broadly known Polish botanist, passed away on 8 August 1994. It is a great loss for the present publication and - on a larger scale - for tropical, and not only tropical, botany. In his memory and with the intention of saving the results of his work we have decided to publish the data existing in Kraków as notes on herbarium labels, separate lists and a card index of species determinations. Text of the present paper was prepared chiefly by Anna Med wee ka-Kornaś (who spent about 2 years in Africa, south of the Sahara) with the technical help of Małgorzata Matyjaszkiewicz who cooperated for a long time with Jan Kornaś and who remembers some facts important for the work; she was involved, among other things, with drawing of diagrams and maps. François Malaisse helped with various information, concerning e.g. description of the study area and location of particular pteridophyte stations. He was assisted by Anne Bodenghien. Some preliminary results of investigations in Upper Katanga, with the information on eleven species new to Zaire (Congo), were already published (Kornaś et al. 1990). In that publication the history of pteridological research in Katanga, or Upper Shaba, may also be found. 1. THE AREA STUDIED AND MAIN TYPES OF VEGETATION 1.1. Location and oro-hydrography Katanga is one of the eight administrative units of the Democratic Republic of Congo in Central Africa and belongs to two phytogeographical regions. Its northern part, with elevation below 850 m is known as Lower Katanga and its 10 ABCDEFGH I JKLMNOPQR 22“ 24" 26“ 28“ 30“ Fig. 1. The location of Upper Katanga (Upper Shaba) on political map of Africa and regional names used in the description of pteridophytes distribution; a - political borders, b - north-western border of Upper Katanga according to the phytogeographical division of Zaire (Congo) (Bamps 1982); c - Katangan Copper Arc southern part (area of presented investigations) is recognized as Upper Katanga and distinguished by higher elevations. It adjoins Angola, Zambia - on the longest section of the border and Tanzania - partly along Lake Tanganyika (Fig. 1). The border line between the northern and southern parts of Katanga is adopted here after Bamps (1982). In Upper Katanga it is necessary to distinguish: a) relatively low territories (below 1000 m a.s.l.), descending to ca 500 m a.s.l. in some river valleys and near the lakes of the Kamalondo - Upemba depression, b) a series of medium high plateaux with altitudes ranging from 800 (1000) m to 1400 (1500) m. On those medium high plateaux the upper courses of some large rivers such as the Lualaba (out of Katanga called Congo), Lufira and Luapula are to be found and c) a series of high plateaux frequently limited by steep escarpments (Fig. 2). Their altitudes reach 2460 m in the north-east, in the Marungu Plateau (called also Marungu Mountains with regard to differentiated relief of its surface). The Ki-bara Plateau near Lake Upemba culminates at 1885 m, and the Kundelungu and Biano Plateaux at 1711 m and 1697 m respectively. In the “Boot of Sakania” Mount Mupwelume reaches altitude of 1893 m. The system of rivers and permanent or seasonal streams is dense and forms a part of the drainage basin of Congo. The southern border of Upper Katanga coincides with the watershed between Congo and Zambezi, two rivers flowing to two different oceans. Several endemic pteridophytes are confined to the area of that watershed (Kornas 1993 a). Many rivers, some of their sections and streams flow in deep ravines. Waterfalls and rapids are numerous, first of all on the escarpments. In their vicinity, the flora of pteridophytes is usually particularly rich. There are, however, also sluggish waters, overflowing seasonally onto flat terraces. Extensive swampy Fig. 2. The relief of Upper Katanga and of adjacent, lower parts of the Republic of Congo (Zaire). The borders as on Fig. 1 12 areas occur mostly at low Luapula (with a set of small lakes), in the basin of the River Lufira and near Lakes Moero and Upemba. 1.2. Climate The climate of Upper Katanga is characteristic of the tropical summer-rain-fall zone, typical for the whole geobotanical Zambezian Region (Walter, Lieth 1960, Fig. 3). The wet season lasts from October - November to early April - mid-May, according to latitude. Mean annual rainfall, following a general pattern, increases from south-east to north-west and amounts to 1100-1400 mm. Some broad valleys present drier conditions with precipitation around 900 mm. Moreover, annual rainfall varies considerably from year to year (820-1780 mm, for instance, for Lubumbashi). There is usually little or even no rain for five to six months. A cool dry season occurs from mid-May to July, whilst a hot dry season is observed in September and October. The mean annual temperature is about 20°C. The existing regional modifications are expressed by the following data: in the mountains near Lake Tanganyika and on the other high plateaux the mean temperatures for July and January are 2-4°C lower than in the adjacent areas. In the south-eastern part (including Lubumbashi region), a second, short, dry season has been observed from time to time (once every four years, on average), mainly in January. The amplitudes of temperatures are larger here than anywhere else. Frost has been recorded in some valleys in the south-eastern part of Katanga, though this remains exceptional (Pain 1978). 1.3. Geology The basic geological structure of Katanga (Fahem 1978 a) consists of old Pre-Cambrian rocks visible in some places on the surface. They have been intensively altered, though to varying degrees, by metamorphism. Among the Lower Pre-Cambrian formations are gneisses (prevailing e.g. on the Muhila Plateau), schists, quarzites and conglomerates. To the Upper Pre-Cambrian formations (classified in the described area as the Katanga System) belong shales, sandstones, dolomites, limestones and quartzites. In the mountains near Tanganyika and on Kibara Plateau are also granites. Some of the rocks mentioned include highly mineralized strata containing heavy metals: copper, cobalt, nickel, radioactive uranium, etc. They occur foremost in the southern region of Upper Katanga, between Kolwezi and Lubumbashi in the so called “Katangan Copper Arc” (or Shinkolobwe Métallogénie Province) and in the Sakania region. Younger deposits are mainly from the Cretaceous or Tertiary Period. Of particular importance among the latter are sands of the Kalahari System (from Miocene and Pliocene), usually unconsolidated. They occur in the western part 13 Fig. 3. Climatic diagrams of seven stations in Upper Katanga and adjacent areas, marked on the map (after Walter, Lieth 1960, changed) 14 of Upper Katanga and on the plateaux: Manika, Biano, Kundelungu and part of Kibara. More recent (from the Quaternary Period) are the colluvial and alluvial deposits which may be found mostly in the river valleys. It should be stressed that in addition to the prevailing siliceous cellular rocks covered by weathered material poor in nutrients there are also in Katanga rare calcareous formations with richer soils. This explains why not only acidophilous but also basiphilous or calciphilous plants can grow there. 1.4. Soils The soils of Upper Katanga are generally poor, not suitable for agriculture. According to the soil map of Zaire (Congo) (Fahem 1978 c) the prevailing part of Katanga is covered by zonal ferralitic soils (sols ferralitiques), occurring mostly on the plateaux and differenciated into ferralsols and arenoferrals. Red-yellow ferralsols are developed on clay or clay-sandy deposits and comprise two main units (cfr. Bridges 1978): a - the leached, very deep soils of the humic tropics, connected with evergreen forests and (in slightly different form e.g. higher content of humus) with some dense riparian and montane forests, and b - the weathered soils of the open forests and savannas. The second type is conditioned mostly by seasonal climate, with rainy and dry periods, typical of Upper Katanga. Changes of soil moisture generate iron concretions present in the soil profiles or as iron pan (or lateric shale) on the soil surface. Some pteridophytes, as will be indicated, grow in such conditions which are unfavorable for many other species, above all for trees. Arenoferrals belong to the “sols lessivés”; they are deeply leached, poor and acid, and developed on sands with low percentage of clay. They can be found under some forests and mostly under savannas. The soils of the Kalahari type sands - pale, loose, and without the iron concretions, which cover the high plateaux - should be classified with this type. Equally important are some azonal or intrazonal soils connected with particular sites. Under seasonally dry or moist conditions, in some depressions and savannas, deep, black, fertile vertisols can be found. There are also alluvial soils - fluvisols, relatively rich in nutrients, with good drainage, occurring mostly under the exuberant, natural vegetation, the boggy (swampy) gley soils developed near some rivers, in depressions, around lakes and springs, and peat-bog soils occurring mainly in the mountains. Quite particular conditions exist on termite mounds, usually constructed from clay-deposits and having more fertile and in some cases drier soil than the adjacent areas. Of importance for ferns and fern allies are also shallow to very shallow initial soils - lithosols, developed on rocks and on slopes where they can be overgrown by open woodlands. Properties of lithosols vary according to the parent material. Of special interest in Upper Katanga are the soils containing heavy metals to which highly interesting and specific vegetation is connected (cfr. pg. 19) 15 1.5. Vegetation The vegetation of Upper Katanga is to a large extent natural or semi-natural consisting of various types of forests, very widespread woodlands, wooded savannas and grasslands. They often form a mosaic in the landscape, caused mainly by very ancient human interference. According to the geobotanical division of Africa (White 1983) the territory of Upper Katanga belongs to the Zambezian Region. This area was formerly considered as part of the Sudano-Zambezian Region which also includes Zambia and some other adjacent countries. Further to the north extends the humid Guineo-Congolian Region, from which some elements penetrate quite far to the south, for instance to Upper Katanga, growing mostly in the “edaphic forests”. Between the two above mentioned regions plant formations of intermediate geographical character have developed (namely savannas described by Mullen-ders 1954). They are specified on the map in Fig. 4 after White (1983) and on a similar map of Fahem (1978 b). Both cited publications are however of a very general character. Various azonal types of vegetation important for pterido-phytes, as e.g. formations along the water courses, are omitted. Therefore in the present description of vegetation, field observations, as well as information from the herbarium labels (unfortunately rather scarce) and from some regional, more detailed publications, including phytosociological maps, are taken into account. Descriptions of vegetation of Upper Katanga are numerous, e.g. Duvig-neaud (1958) and Schmitz (1963). Some of them concern particular regions e.g. Katangan High Plateaux (short paper of Lisowski et al. 1971), Kundelugu Plateau (Malaisse 1975), Lowlands of the River Lufira (Streel 1963), Plain of Lubumbashi (Schmitz 1971), Upemba National Park (VanMeel 1966) and Ka-niama situated beyond, but not far from the area under consideration (Mullen-ders 1954). There are also publications about selected vegetation types: forests (Lebrun, Gilbert 1954), muhulu forest (Schmitz 1962, Malaisse 1984), miombo woodland (e.g. Malaisse 1997), aquatic and swamp vegetation (e.g. Lisowski, Malaisse 1989), about flora and vegetation on the termite mounds (Malaisse 1978) and metalliferous soils (e.g. Duvigneaud 1958, 1959, Duvigneaud, De-naeyer-de Smet 1963, Brooks et al. 1985). A large part of those studies contains phytosociological descriptions of plant associations. The review presented here is very simplified. It contains however some French definitions used in literature and on herbarium labels. Vegetation types mentioned below can be considered as the most important in Upper Katanga. Montane (Afromontane) formations - they occur on the highest elevations, mostly on the Marungu Plateau (or Mountains) and need to be further studied. In all probability they are divided according to altitude into: 16 Fig. 4. Main vegetation types in Upper Katanga and lower, adjacent part of Congo; 1 - montane (Afromontane) vegetation, 2 - Guineo-Congolian rain forest of dryer type, 3 - mosaic of lowland rain forest and secondary grassland (Guineo-Congolian), 4 - wetter Zambezian miombo woodland, 5 - mosaic of Zambezian dry evergreen forest “muhulu” and wetter miombo woodland, 6 - mosaic of wetter Zambesian woodland and secondary grassland, 7 - secondary wooded grassland with Acacia polyacantlia, 8 - edaphic and secondary grassland on Kalahari sands, 9 - herbaceous swamp and aquatic vegetation. After White (1983), slightly changed (for further explanation see pg. 15-19) a) Montane forests at altitudes above 1600 m, conditioned by higher humidity (e.g. mist concentration). They have a character of dense montane forests - forêts denses montagnardes - with tree ferns, climbers, epiphytic ferns, mosses, lichens, etc. (as on the escarpments near Lake Tanganyika) or of forests with Acacia abyssinica, many shrubs and tall herbs. In some places, mostly near high springs, the giant Senecio and Lobelia can be found. b) Sclerophyllous scrub (thicket) - maquis sclérophylle - after Duvio-neaud (1958) occurs above tree limit (above 2000 m). Very typical here is the 17 presence of Ericaceae, Proteaceae, possibly Juniperus procera (in one locality). Higher up are alpine grasslands limited to some summits, including numerous species of Helichrysum. c) Montane peat bog - tourbière - is also found at high altitudes and composed of Sphagnum planifolium with various species of Drosera, Cyperus, etc. Group of dense (not montane, or low montane) forests, conditioned edaph-ically - forêts édaphiques. Typical of undisturbed stands are several layers of vegetation and closed canopy. These forests, evergreen or semi-evergreen, are in general connected with moist and wet sites (in many cases with high ground water level) and soils relatively rich in nutrients. They offer very suitable habitats for pteridophytes and are differenciated according to site conditions into: a) Riparian forest (riverine forest, gallery forest) - named on the labels forêt galerie or galerie forestière - occurs near waterfalls, along larger streams and rivers, mostly on flat valley bottoms, on well drained or slightly swampy soils, usually (but not exclusively) on flooded terraces. Occurrence of trees as e.g. species of Pandanus, Raphia and Ficus, of Phoenix reclinata and Syzygium guineense depends as well on the distance from the water courses. Among climbers e.g. Mezoneuron was noted. This type of forests is often disturbed by natural factors (floods) and human exploitation, which open gaps suitable for some light demanding species. b) Ravine dense forest - forêt dense de ravin (often mentioned in the herbarium collection) - occurs in ravines incised mostly in the escarpments of the plateaux. As evergreen formation it may be connected with rivers and perennial or seasonal streams, or even with some wet and shady slopes. To the species occurring here belong e.g. Parinari excelsa, Podocarpus milanjianus (Conife-rae) and tree-fern Cyathea thomsonii. c) Narrow fringing forest (fringing forest) - frange forestière étroite - this formation of low, evergreen trees and shrubs, rich in epiphytic ferns forms narrow strips along streams flowing in very shallow gullies, incised in the surface of the plateaux. The floristic composition of fringing forests is slightly different in middle altitudes than in higher sites above 1450 m, where e.g. Agauria sa-licifolia grows and more often Osmunda regalis can be found. d) Swamp forest - forêt marécageuse, “mushitu” - connected with weakly drained soils forms stands in swampy places, in depressions, on alluvial terraces where the complexes with riparian forests can form. Trees, e.g. Mitragyna stipu-losa are tall, with buttresses, stilt roots and sometimes with pneumatorhize. On high plateaux the swamp forest occurs mainly around springs and is distinguished by the moss Sphagnum planifolium. Dense, relatively dry forest - forêt dense sèche or forêt dense non édaphique - “muhulu”. This forest is usually semi-evegreen, with presence of deciduous species in the tree layer and with evergreen undergrowth. It has a closed canopy, rather rich floristic composition and usually forms islands among more 18 open formations. “Muhulu” is considered by some authors as climax on large territories of Upper Katanga disturbed by human influence. Two types of this forests can be distinguished: plots (communities) with Entandrophragma dele-voyi on deep, clay (argileux) soils and plots with Cryptosepalum exfoliatum and Marquesia acuminata on sandy soils. Open woodlands of zambezian type — forêts claires zambéziennes — here belong forests with trees in single storey and light, open canopy, deciduous or semi-deciduous. The ground layer is often distinguished by a high percentage of grasses. The soil, poor in nutrients, can be very stony, chiefly on some summits and slopes. Most widely spread (first of all on medium high plateaux) are open woodlands “miombo” with domination of various species of Brachystegia (e.g. B. spiciformis), Julbernardia and Isoberlinia. In more favorable conditions taller trees as Marquesia macroura can be found. Bushland and thicket - bosquet - may occur on transgression between various vegetation types as e.g. belts with Uapaca robynsi between “cupriferous steppes” and forests. Often bushlands and thickets are formed by anthropogenic impact upon the vegetation. Wooded savannas or wooded grasslands - here belong savane boisée and savane arborée - both with trees more than 8 m tall; there is also savane arbus-tive with shrubs. In general tree-crowns cover 30-60% of the area for “savane boisée”, less for “savane arborée”. From a geographical point of view can be distinguished: “wooded savanna of Guinean type” occurring mostly in northern and lower parts of Upper Katanga and more xeric “Zambezian Savanna” (with e.g. Borassus flabelliformis subsp. aethiopicum), which covers open spaces in the range of “miombo” woodland. Both types of formations are usually due to human impact, first of all by the use of fire, resulting in the expansion of Acacia polyacantha. Tree savannas can exist however owing to natural, local factors such as poor, dry soil or floods maintaining their alluvial types. Open savannas or edaphic and secondary short grasslands - les savanes steppiques ou steppes (“dilungu”). Particularly worthy of attention are formations on Kalahari sands, seasonally very dry, covering the high plateaux of Katanga. They are formed by short cespitose grasses of Poaceae and by Cyper-aceae apart from which there are some dicotyledonous species of Fabaceae, Lentibulariaceae, herbaceous Lobeliaceae, etc. Especially interesting here is the ecological group of fire resistant “geoxylic suffrutices” such as Parinari and Syzygium. Short grassland (steppe) also cover the main part of hills with soils containing heavy metals. In some seasonally wet places on the plateaux “steppe humide" can be found. Moist meadow (“dembo”, “dambo”) - exists on sites with permanently seeping water or in seasonally wet depressions and are composed of grasses, sedges, with a high share of dicotyledonous plants and some orchids. On some places in that formation groups of palms Raphia can occur. 19 Swamp herbaceous vegetation - végétation marécageuse - may be found in depressions, on the banks of the lakes and larger rivers and consists of communities with grasses Leersia and Vossia, sedges e.g. Cyperus papyrus, of reeds Phragmites mauritianus and Typha capensis. Swamp herbaceous vegetation usually forms a complex with aquatic plant formations. Aquatic vegetation - generally develops in stagnant waters and sluggish river arms and is composed of submerged or floating plants, e.g. Nymphaea spp. and Pistia stratiotes; it is the habitat of fems Marsilea and Azolla. Vegetation of high termite mounds (or termitaria) - végétation des hautes termitières - is not uniform and depends upon diversified soil and light conditions as well as termite activity. Plant communities there are usually different from the surroundings and composed of various ecological groups of species (among others xerophytes and some ferns). They form particular flore or végétation termitophile (Malaisse 1978). Vegetation of rocks and rock crevices - has the share of lithophytic ferns - calciphilous as e.g. Adiantum capillus-veneris and Elaphoglossum chevalieri, acidophilous (or indifferent) as e.g. Arthropteris orientalis and Hypodematium crenatum. The majority of lithophytic pteridophytes was found in Upper Katanga on humic and shaded rocks, situated in the forests, near waterfalls and rivers. The high plateaux escarpments provide rocky situations especially suitable for various ferns. To about 30 heavy metal tolerant species belong e.g. Actiniopteris kornasii, Pellaea pectiniformis and Mohria lepigera. That group occurs mainly in the “Katangan Copper Arc”, on the rock outcrops among open vegetation “savonnes stepiques” (Brooks et al. 1985, Malaisse et al. 1994). Worthy of attention are the diagrams illustrating the spatial distribution of various vegetation types in “catenas” e.g. on Kundelungu Plateau and in Luanza Valley (Malaisse 1975) or in the “Katangan Copper Arc” (Duvigneaud 1958, Malaisse et al. 1994). In the last area numerous plant communities analogical to those occurring in other parts of Katanga were found; their particular feature is however the presence of rare, endemic species. Among the important anthropogenic factors influencing the vegetation in Upper Katanga can be mentioned: timber exploitation for charcoal production, common use of fire to eliminate trees, shrubs as well as high grasses, the raising of cattle which show the greatest concentration on the Biano and Kundelungu Plateaux and in the lower montane regions near Tanganyika, belonging mainly to the Plateau Marungu. Of importance also is agriculture, but cultivated fields cover in general less than 1 ha per 1 sq. km; apart from some plantations there are usually small fields which are abandoned after some years. Then the land is usually overgrown by grasses; on some disturbed areas Pteridium aquilinum can dominate. Exploitation of mineral resources in the southern part of Katanga takes place mostly in open quarries. Waste heaps connected with it form special sites for plants, among others pteridophytes. The existence of industry generates 20 some air pollution. There are also dams and man made lakes, e.g. on the River Lufira. It should be mentioned that a large part of the Kibara Plateau and northern part of Biano Plateau with adjacent lowlands are included in the Upemba National Park (950 000 ha), and that an another National Park has been established on the southern part of the Kundelungu Plateau, with adjacent foothills (22 000 ha). 2. Methods, material and scope of information Pteridophytes were collected by A. Bodenghien and F. Malaisse during various vegetation seasons and from more than 300 stations (Fig. 5), which were chosen randomly, mostly near the roads and in particularly interesting sites as 22“ 24° 26“ 28“ 30“ Fig. 5. Stations of pteridophyte collections by A. Bodenghien and F. Malaisse; a - presented on the dot map, b - presented by the grid method 21 the vicinity of waterfalls; later on these data were transferred into information unified according to the grid method (see below). The herbarium collection numbers about 5 000 voucher specimens. The materials - representing all taxa - were sent to KRA where their determination was made, or checked, by Jan Kornas. He also worked out some taxonomic revisions and noted critical remarks, which are included in the present text. As a material for comparison Kornas used chiefly his own collections from Zambia, Northern Nigeria, Tanzania and some parts of Cameroon as well as the collection of Kazimierz Nowak from Rwanda determined also by Kornas. He had also an insight into other Herbaria, e.g. BR, GENT, K (Kew), and cooperated with pteridologists in numerous countries. In the present study, apart from the collection of A. Bodenghien and F. Ma-laisse housed in KRA, materials of other botanists, mainly of Stanislaw Lisowski in the same herbarium as well as data from floristic publications, as e.g. fascicles of the “Flore d’Afrique Centrale” were taken into account. Specimens of S. Lisowski from POZG (formerly POZ) are quoted after Kornas (1979). Owing to those citations data from other herbaria (not only KRA) are also included. The important part of African fern collections in BR and BRLU, e.g. herbarium of Paul Duvigneaud, is waiting, however, still for determination. There are various systems of pteridophyte classification (e.g. Holttum 1974), discussed in relation to an older publication by Pichi Sermolli (1973). We adopted here the classification of Kramer and Green (1990), different from those used e.g. in the study concernig the neighbouring Zambia (Kornas 1979), but more recent (Table 1). A little bit later, conceptions of Kramer and Green, were however changed in some details in the paper of Johns (1991). In the list presented fern families are arranged alphabetically, and so are the genera in frames of families and species within the genera. A few specimens left by J. Kornas with preliminary or doubtful determinations are also mentioned (usually with a mark “cfr.”). They are grouped separately when doubtful determinations concern relatively numerous specimens and are in part supplemented by some taxonomic remarks. It concerns first of all lower, intraspecific units, deserving of further studies. Nomenclature of species is given according to the decisions of Kornas and coincides as a rule with the recent literature. The synonyms are introduced selectively, mainly when they are useful for orientation in former, quoted litterature. For particular taxa short information about frequency, general distribution in Upper Katanga and ranges of altitude is given. Frequency of species, or lower units, is estimated according to the number of stations, in the following scale: 1-4 - very rare, 21-30 - common, 5-10 - rare, above 30 - very common. 11-20 - quite common, 22 Table 1 Differences in pteridophyte families classification GENERA FAMILIES- according to the quoted authors (in order of the Kramer & Green Crabbe, Jermy Johns present paper) (eds.) 1990' & Mickel 1975” Schelpe 1970 & Edwards 1991 (in Johns 1991) Athyrium Dryopteridaceae Aspleniaceae Athyriaceae Woodsiaceae Ctenitis Dryopteridaceae Aspleniaceae Aspidiaceae Dryopteridaceae Diplazium Dryopteridaceae Aspleniaceae Athyriaceae Woodsiaceae Dryopteris Dryopteridaceae Aspleniaceae Aspidiaceae Dryopteridaceae Hypodematium Dryopteridaceae Aspleniaceae Aspidiaceae Dryopteridaceae Polystichum Dryopteridaceae Aspleniaceae Aspidiaceae Dryopteridaceae Tectaria Dryopteridaceae Aspleniaceae Aspidiaceae Dryopteridaceae Bolbitis Lomariopsidaceae Aspleniaceae Lomariopsidaceae Lomariopsidaceae Elaphoglossum Lomariopsidaceae Aspleniaceae Lomariopsidaceae Lomariopsidaceae Lomariopsis Lomariopsidaceae Aspleniaceae Lomariopsidaceae Lomariopsidaceae Nephrolepis Nephrolepidaceae Davalliaceae Davalliaceae Oleandraceae Arthropteris Oleandraceae Davalliaceae Davalliaceae Oleandraceae Oleandra Oleandraceae Davalliaceae Davalliaceae Oleandraceae Loxogramme Polypodiaceae Grammitidaceae Polypodiaceae Polypodiaceae Actiniopteris Pteridaceae Adiantaceae Adiantaceae Actiniopteridaceae Adiantum Pteridaceae Adiantaceae Adiantaceae Adiantaceae Aspidotis Pteridaceae Adiantaceae Adiantaceae Adiantaceae Cheilanthes Pteridaceae Adiantaceae Adiantaceae Adiantaceae Ceratopteris Pteridaceae Parkeriaceae Adiantaceae Parkeriaceae Coniogramme Pteridaceae Adiantaceae Adiantaceae Adiantaceae Doryopteris Pteridaceae Adiantaceae Adiantaceae Adiantaceae Pellaea Pteridaceae Adiantaceae Adiantaceae Adiantaceae Pityrogramma Pteridaceae Adiantaceae Adiantaceae Adiantaceae Pteris Pteridaceae Adiantaceae Adiantaceae Pteridaceae Vittaria Vittariaceae Adiantaceae Vittariaceae Vittariaceae * Classification adopted by the present authors and by KornaS et al. (1993) in pteridophyte flora of Rwanda ** Classification adopted by KornaS (1979) in pteridophyte flora of Zambia Descriptions of distribution are based on geographical names indicated in Fig. 1. Altitude mentioned for particular stations concerns in some cases the sites of various collections, not strictly the occurrence of a particular fern or fern allies. It is so when a relatively great altitudinal range (e.g. of 100 m) is indi 23 cated. A brief description of habitats is based mainly on the herbarium labels. More detailed ecological descriptions and other remarks can be found for a prevailing number of species growing in Upper Katanga in the pteridological study from Zambia - Kornas (1979), on the pages mentioned by present authors. The location of stations of particular taxa is indicated in the present text with the use of a grid method. The squares marked by letters and numbers (cfr. e.g. Fig. 5) are always mentioned in order from West to East and from North to South. The maps of taxa distribution (Figs 6-202) are constructed according to the same grid, or cartogram method (only one mark in each square indicating the occurrence of a given species in the related area). The distance between the lines of the grid relates to 0.5° of geographical latitude and longitude. The location of stations was assigned according to the information of Belgian botanists, index of collecting places compiled by Bamps (1982), and some physical, detailed maps. Stations situated just on the lines of the grid are indicated in squares adjacent to them on the right or below. For synonyms of local topographic names and information about some tributary rivers - see Appendix. Abbreviations The herbarium abbreviations are those proposed by Holmgren et al. (1990). Abbreviations of collector names: B&M - Bodenghien and Malaisse Rob. - Robyns Lew. - Lewalle Robbr. - Robbrecht Lis. - Lisowski Schj. - Schaijes Luk. - Lukuesa Schm. - Schmitz Mai. - Malaisse Sym. - Symoens Others abbreviations: alt. - altitude Rap. - Rapids confl. - confluence of rivers Rav., rav. - Ravine, ravine env. - in environs of R. - River Esc. - Escarpment Spr., spr. - Spring, spring ibid. - ibidem Stm., stm. - Stream, stream Miss. - Mission terr. - territory Mt. - Mountain trib. - tributary river Natl. - National Val., val. - Valley, valley Note. Station location: e.g. “Kolwezi - Kamina 52 km” means 52 km from Kolwezi to Kamina, on the road between these stations. Numbers in the text with meters (e.g. 1000 m) means altitude. 24 3. List of species Note. All voucher specimens indicated below by numbers are housed in KRA Herbarium. The asterisk (*) at some numbers of Figs 6-202 means the distribution maps concerning the given species together with its lower units, subspecies or varieties. Psilotaceae Psilotum nudum (L.) P. Beauv. Distribution. Fig. 6. Rare, Kundelungu Plateau - its lower parts and periphery, region of Lubumbashi, alt. 1130-1225 m (after 5 records), 8 stations. К 13: Kimilolo R„ 1225 m, B&M 120, Lubumbashi (Lawalree 1969 d, Kornas 1979), 11 km from Lubumbashi, 1220 m (Schelpe 1973, Kornas 1979); L 10: Luanga R., “contreforts” of Kundelungu, 1130 m (KornaS 1979), Luanza Basin (Malaisse 1975, KORNAS 1979); L 12: Luiswishi R„ 1200 m, B&M 183, 196; M 10: near Kabia-shia (Lawalree 1969 d, Kornas 1979), 4 km WNW from Kabiashia, ИЗО m (Schelpe 1973). Habitat. Terrestrial or epiphytic on lower parts of tree trunks, in riparian forests, near rivers and waterfalls, occasionally in rock crevices. Zambia - Kornas (1979): 26. Lycopodiaceae Taxonomic remark. Classification of Lycopodiaceae follows the system adopted by Kornas (1979) and also e.g. by Burrows (1990) and Jacobsen (1983); more recent names are mentioned as synonyms. Lycopodium carolinianum L. (= Lycopodiella carolinianum (L.) Pic. Ser.) Taxonomic remark. A polymorphic taxon. A part of the herbarium material in KRA was left by J. Kornas without varietal names. It concerns tuberless voucher specimens, determined by collectors and authors of former, quoted publications as var. affine (or Lycopodiella affinis) - those published determinations are respected in the text below. Specimens in KRA, with tubers and tuber initials, were distinguished by Kornas as var. tuberosum. He stressed however (Kornas 1979) that separation of the varieties “affine" and “tuberosum” seems to be extremely doubtful. Information about the general distribution and habitat concerns therefore jointly Lycopodium carolinianum s. 1. and its two varieties. 25 Distribution. Fig. 7*. Very common, region of Lualaba River, plateaux: Manika, Biano, Kundelungu, Muhila and Marungu, south-eastern border of “Sa-kania Boot”, alt. 1215-1900 m (after 29 records), 35 stations. L. carolinianum s. 1. G 11: Manika Plateau, 1450 m, Mai. 8737; H 10: Katentania, Lis. Mai. & Sym. 13696', 110: Kipao Spr., 1630 m, B&M555; L10: Kapala R., 2 km N from Luishi, 1645 m, B&M 1020; P15: Malebwe R., Mupala, 1210 m, B&M 2244; P16: Chinika R., 1500 m, B&M 2650, Fibulukutu Spr., 1450 m, B&M 2588, Kasamba R., 1250 m, B&M 2727, Lusemfwa R., 1500 m, B&M 2630, Mabondo R., between Ifuna and Malashi, 1290 m, B&M 2318, Miloso R., Mupala, 1220 m, B&M 2750, Nyamanda Spr., 1530 m, B&M 2669. L. carolinianum var. affine (Bory) Schelpe (= Lycopodiella affinis (Bory) Pic. Ser.) G 10: Munanga, 90 km N from Kolwezi (Lawalree 1989); G 11: S from Kolwezi (Lawalree 1989); G 12:40 km S from Sakabinda (Lawalree 1989); 110: Biano-Gare, 1580 m (Schelpe 1973, Lawalree 1989); L 10: Katshupa (Lawalree 1989), 6 km NNW from Poste de Katshupa, 1690 m (Schelpe 1973), Luishi (LawalrBe 1989), Lut-shipuka Spr., 1600 m, Lis. 10087, (Kornas 1979, Lawalree 1989), 12 km WNW from Msipashi, 1700 m (Schelpe 1973), 15 km SW from Msipashi, 1660 m (SCHELPE 1973); L 11: Kalembe R., 1550 m (KORNAS 1979); O 4: Lukanda swamp, 1470 m, B&M 1788, (Lawalree 1989); P5: Pepa, 1900 m (Kornas 1979, Lawalree 1989). L. carolinianum var. tuberosum (Welw. et A. Braun ex Kuhn) Nessel L 10: 6 km N from Poste de Katshupa, 1680 m (KORNAS 1979), Luanza Basin (Ma-laisse 1975, Kornas 1979), Luanza Spr., 1690 m B&M 1002; L 11: Lutshipuka Spr., 1670 m, B&M 966; O 4: Muizia R„ 1450 m, B&M 740; P 15: Kabeleshi R., 1230 m, B&M 2225, (Lawalree 1989 - as Lycopodiella affinis); P 16: Akapemfia R., 1500 m, B&M 2610, Katuba R„ 1215 m, B&M 2256, Kasala R„ Mupala, 1245 m, B&M 2270, Umusangashi R., 1305 m, B&M 2289, Umwimbwa R., 1300 m, B&M 2280. Habitat. Terrestrial, in humid and wet sites, near rivers and montane springs, in swamps, peat bogs, on boggy slopes and in moist meadows, at the edges of swamp forests in alluvial savannas (relatively often), occasionally on sand, in full sunlight or in very slight shade. Zambia - Kornas (1979): 26 - L. carolinianum var. affine and var. tuberosum. Lycopodium cernuum L. (= Lycopodiella cemua (L.) Pic. Ser.) Distribution. Fig. 8. Very common, western part of Upper Katanga at Di-lolo, region of upper Lualaba River, plateaux: Manika, Kibara (Park National Upemba), Biano, Kundelungu, plateau near Tanganyika, region of Luvua River, of Lubumbashi, Sakania and south-eastern border of “Sakania Boot”, alt. 810— 1840 m (after 55 records), 64 stations. 26 A 11: Dilolo (Lawalree 1989); G 10: Kolwezi - Kamina 52 km (Lawalree 1989), Mushinji R., Kolwezi - Kyamasumba 70 km, 1010 m, B&M 2150, (Lawalree 1989), Mutendele R., 810 m, B&M 370; G 11: Kisote, 1480 m (Kornas 1979, Lawalree 1989), Manika Plateau, 1450 m, Mai. 8791, 8737 bis, Potopoto trib., Kolwezi - Muso-katanda 30 km, 1425 m, B&M 340; H 10: Katentania, 1370 m (SCHELPE 1973, Lawalree 1989), Katentania R., 2 km from “poste de Katentania”, 1500 m, B&M 704; I 10: Kansenia (Lawalr&e 1989), Kipao R„ 1590 m, B&M 647, Kisamoya R., 1500 m, B&M 632, Menge R., near Quaglia Farm, 1575 m, B&M 569, Mulamba Falls, 1510 m, B&M 132; J 7: Kabwe, Upemba Natl. Park (Lawalree 1989); J 8: Kaziba - ibid. (Lawalree 1989); K 6: Mulumbi Esc., Mitwaba - Manono 52 km, 950 m, B&M 478; K 7: Mitwaba - Manono 11 km (Schelpe 1973, Lawalree 1989), ibid. Bana Bankasi Hill, B&M 456; K 13: Kisanga (Lawalree 1989), Kipopo, 1230 m (Schelpe 1973, Lawalree 1989), 15 km SE from Lubumbashi, 1150 m (SCHELPE 1973); L 8: Kimitondo Stm., Kibwe wa Sanga, 1730 m, B&M 1040, Kwamba R., 1650 m, B&M 1045, Mwanza R., 1640 m, B&M 1030; L 10: Kapala R., 2 km N from Luishi, 1645 m, B&M 1025, Lualala, 1660 m (Kornas 1979), Luanza Basin (Malaisse 1975, Kornas 1979), Luanza R., 6 km NNW from Poste de Katshupa, 1690 m (Schelpe 1973), Luanza Spr., 1690 m, B&M 987, (Lawalree 1989), Nungwe trib., 1670 m, B&M 1071; L 11: Kalembe R„ 1550 m, Lis. Mai. & Sym. 12769, (KORNAS 1979), Luanza Vai., 6 km N from Katshupa, 1680 m, B&M 258; LutshipukaR. (Lawalree 1989); L12: Ponde Stm., Mwenge, 1090 m, B&M 950; L 13: Munama R., Kasamba, 1220 m, B&M 1587; L 14: Kinsenda, 1330 m, B&M 2083, (Lawalree 1989); N 5: Mukui R„ 1300-1360 m, B&M 3570; N 15: Kalumbwe R„ 1230 m, B&M 1977, Lukusashi R„ 1230 m, B&M 1965; O 4: Kamabale R„ 1350-1420 m, B&M 1639, Kamakubi R., 1500 m, B&M 1737, Katomia R„ 1400 m, B&M 2957, Kilumbi R., Kilenge Rap., 1450-1500 m, B&M 1301, Kipanda R., near Kasimba, 1550-1575 m, B&M 1194, (Lawalree 1989), Kitu R„ 1500 m, B&M 1745, Laula Rav., 1350-1400 m, B&M 1258, Lukonagulwa Stm., 1460 m, B&M 2815, Mazala Stm., 1700 m, B&M 1729, Mpa Stm., 1420 m, B&M 2931, Muizia R., 1450 m, B&M 739, Musofwa R., near the dam, 2 km from Poste Musofwa, 1350 m, Lis. 10105; P 4: Mulobozi R., Kapupo Fall, 1020-1100 m, B&M 2762; P 5: Kasia R., Kieluzi - Mwela 40 km, 1780 m, B&M 1440, (Lawalree 1989), Lufuko R„ near Pepa, 1790-1840 m, B&M 1493; P15: Kabeleshi R„ 1230 m, B&M 2226, Malebwe R„ 1190 m, B&M 2243; P 16: Akapemfia R., 1500 m, B&M 2613, Bolelo R., Serenje, 1370 m (SCHELPE 1973), Chinika R., 1500 in, B&M 2654, Kamakolongo R., 1440 m, B&M 2480, Miloso R., Mupala, 1220 m, B&M 2748, Mubwe R., 1350 m, B&M 2411, Nyamanda R., 1290 m, B&M 2389, Tchitwichan-sofu R., 1440 m, B&M 2445. Habitat. Terrestrial, on river banks, in fringing, riparian and swamp forests and at their edges, in open forests e.g. on moist sites between the rocks, in alluvial savannas mostly on sandy places; in shade and in full sunlight. Zambia - Kornas (1979): 28. Lycopodium clavatum L. Taxonomic remark. Voucher specimens in KRA are left by J. Komas without varietal names, even when they are indicated by some collectors. Citations after Lawalree (1989) concern also L. clavatum s. 1. In a publication about 27 Zambia (Kornas 1979) and in some other papers var. inflexum (= var. borboni-cum) was however distinguished. Therefore that variety is quoted below separately (taking into consideration also such voucher specimens for which var. inflexum was indicated at least by one author only). Information about the general distribution and habitat of L. clavatum in Upper Katanga leaves out the varietal distinction. According to Kornas (l.c.) the taxonomic status of L. clavatum in Africa requires further critical study. Distribution. Fig. 9*. Quite common, Biano Plateau and more often - region of Luvua River and Muhila Plateau, also Marungu Plateau, alt. 1290-2200 m (after 17 records), 17 stations. L. clavatum s. 1. I 10: Kisamoya R., 1500 m, B&M 638, (Lawalree 1989); N 5: Kibondo R., 1335 m, B&M 3029, Lualashi R., 1300-1350 m, B&M 3054, Mukui R„ 1300-1360 m, B&M 3572, Mukui trib., 1290-1360 m, B&M 3612, 3640, 3668; O 4: Katomia R„ 1400 m, B&M 2958, Kipanda R., 1550-1575 m, B&M 1193, (Lawalree 1989), Libanda R., 1450 m, B&M 2896, Lukonagulwa Stm., 1460 m, B&M 2816, Mpa Stm., 1420 m, B&M 2932; P5: Kasia R„ Kieluzi - Mwela 40 km, 1780 m, B&M 1441, (Lawalree 1989), Lufuko R., near Pepa, 1790-1840 m, B&M 1494, (Lawalree 1989), Mufufu R., 1900 m, B&M 1575. L. clavatum var. inflexum (P. Beauv.) Spring (= L. clavatum var. borbonicum Bory) I 10: Biano II, 1600 m, Lis. Mai. & Sym. 13694, (LISOWSKI et al. 1973, Schelpe 1973, Kornas 1979, Lawalree 1989), Grelco, 1600 m (Lisowski et al. 1973, Kornas 1979, LAWALREE 1989); P5: Lunanga Spr., 2200 m (LISOWSKI et al. 1973), Mulombozi Spr., near Kasiki, 2200 m (Lisowski et al. 1973, Lawalree 1989) - two last stations are mentioned by SCHELPE (1973) generally as Marungu Plateau. Habitat. Terrestrial, at the edges of riparian forests, in some disturbed forests and thickets, near montane springs, in ravines; on moist sand and rocks. Zambia - Kornas (1979): 29 - var. inflexum only. Lycopodium dacrydioides Bak. (= Huperzia dacrydioides (Bak.) Pic. Ser.) Note. Voucher specimens in KRA were determined by J. Kornas as L. dacrydioides and this distinction is adopted here in spite of the fact that a number B&M 1486 is regared by Lawalree (1989) as an other species, Huperzia mild-braedii (Herter) Pic. Ser. Distribution. Fig. 10. Very rare, Marungu Plateau, 2 stations. P5: Lufuko R„ 7 km from Pepa, 1790-1840 m, B&M 1486, (LAWALREE 1989 as Huperzia mildbraedii (Herter) Pic. Ser.), Malonge R., 1800 m, B&M 4147. Habitat. Epiphytic or lithophytic (with pendulous stems), in riparian forest on steep banks and on the rocks in river rapids and waterfalls. Kornas (1979): 30, in Zambia not noted, information from Nyika Escarpment, Malawi. 28 Lycopodium gnidioides L. f. (= Huperzia mildbraedii (Herter) Pic. Ser.) Note. The occurrence of L. gnidioides in Upper Katanga was announced by Lisowski et al. (1973) and Schelpe (1973). Information is based on specimens not seen by Kornas and mentioned later under the name H. mildbraedii by La-walrée (1989). Such determination, cited below, is perhaps not definitive (as in the case of revised specimen classified at present as L. dacrydioides - see above). Further taxonomic studies are therfore needed. Distribution. Fig. 11. Very rare, Muhila Plateau, 1 station. О 4: rav. of Laula R., 1350 m (LISOWSKI et al. 1973, SCHELPE 1973, Lawalrée 1989 - as Huperzia mildbraedii). Habitat. Epiphyte pendulous or erect, growing usually in montane forests. Station at Laula R. is in all probability connected with local cool conditions existing in the ravine, near waterfall. Kornas (1979): 30, in Zambia not noted, reported from Nyika Escarpment in Malawi. Selaginellaceae Selaginella abyssinica Spring (= S. goudotiana Spring var. abyssinica (Spring) Bizzarri) Distribution. Fig. 12. Very common - region of rivers Lualaba, Lufira, Luvua and Lubule, plateaux: Biano (numerous data), Kibara, Kundelungu and some of their surroundings, Muhila and Marungu, sites near Lake Tanganyika, Plain of Lubumbashi, alt. 950-2000 m (after 54 records), 73 stations. G 10: Luilu trib., near Kolwezi, 1130 m, B&M 355; H 9: env. Lubudi (Bizzarri 1985); H 10: Kalule Sud R., 10 km from Lubudi, 1260 m (SCHELPE 1973), Katentania R„ 1500 m, B&M 705; H 11: Kolwezi - Manfwe 12 km, 1350 m, B&M 2025; I 9: Dipera Falls, 1220 m, B&M 423, Kayo Falls, 1250-1310 m, B&M 420, 422, Kia Mt., Upemba Natl. Park (BIZZARRI 1985), Lubudi - Nguba 14 km, 1450 m, B&M 440; 110: Guie R., 1450 m, B&M 547, 548, 549, Kadiva Falls upstream of Kansenia, 1300 m, B&M 607, Kalule Sud R., 1500 m, B&M 620, Kisamoya R., 1500 m, B&M 637, Mu-lamba Falls env. Kansenia, 1250 m (Kornas 1979), Mulamba Falls, Kapiri, 1350 m (Schelpe 1973, Bizzarri 1985), Mulamba R., 1460 m, B&M 671, Mulandakipindi Stm., 1450 m, B&M 697, Pande Fall upstream of Kansenia, 1300 m, B&M 594, 595, 597, (Bizzarri 1985); I 11: Fungurume 1220 m (Schelpe 1973, Kornas 1979, Bizzarri 1985), Fungurume “Hill V”, 1130 m, B&M 2067, Kimisombo Stm., Kakanda - Kambove 9 km, 1430 m, B&M 883, Kisikimalwa Mt., Fungurume - Nguba 9 km, 1220 m, B&M 2133, Luita, 1520-1550 m, B&M 887, Luita trib., 1480 m, B&M 895, Tshilongo Stm., 1350 m, B&M 325; J 7: Katongo R., Upemba Natl. Park (Bizzarri 1985); J 11: Dilomba R. (Bizzarri 1985), 5 km N from Kakontwe, Mai. 11721, (Bizzarri 1985), 6 km SSE from Kambove, 1360 m, B&M 838, Kanduba R., 8 km W from Kakontwe, 1330 m, B&M 833, Likasi, 1130-1150 m (SCHELPE 1973, KORNAS 1979, 29 BlZZARRI 1985); K 6: Mulumbi Esc., Mitwaba - Manono 52 km, 950 m, B&M 477; K 11: Mwadingusha, 1000 m (SCHELPE 1973, BlZZARRI 1985), Namuntamba Mt., 1180 m, B&M 2119; K 12: Kamianga Stm., 5 km N from Shilatembo, 1200 m, B&M 286, 321, 322, Luafi Stm., Lubumbashi - Likasi 70 km, 1230 m, B&M 304; K 13: Kafubu R., Ki-sanga, 1235 m, B&M 792, Karavia, 16 km W from Lubumbashi, 1150 m, Mai. & Robbr. 2090, Kasapa R. (BlZZARRI 1985), Kisanga near Lubumbashi, 1200 m (Schelpe 1973, BlZZARRI 1985), Lubumbashi R. (BlZZARRI 1985), Lupoto (BlZZARRI 1985); L 8: Mu-pungwe Falls, Sampwe (BlZZARRI 1985); L 10: Kaloba (BlZZARRI 1985), Luanza Bassin (MALAISSE 1975, KORNAS 1979), Munyankara Stm., env. Gombela, 1060 m, B&M 391; L11: Lukafu (BlZZARRI 1985), Lutshipuka Falls, 1550 m, B&M 231, 237, Masansa Fall, 1500 m, B&M 280; L 12: Luiswishi, 1208 m, B&M 32, 299, 773; L 13: Katuba R., Munkeo, 1220 m, B&M 519; L 14: Kasumbalesa Mt., 1380-1450 m, B&M 2075; M 6: env. Kisele, B&M 82; M 10: Luanza Vai., W from Kabiashia, 1350 m (Schelpe 1973, BlZZARRI 1985), Msipashi R., env. Kinika, 1180 m (KornaS 1979, BlZZARRI 1985); M 13: Lubembe, Mokambo - Kasumbalesa road (Schelpe 1973, BlZZARRI 1985); N 7: Mulonde R., Kanfwa Fall, Kyankalamu - Pweto 48 km, 1030 m, B&M 1378; O 1: Ten-dele R., Tugulu (Demaret 1955), between Tugulu and Luama R. (Demaret 1955); O 4: Kapona Mt. (BlZZARRI 1985), Kyula Mt., env. Kansimba, 1680 m, B&M 750, (BlZZARRI 1985), Kasenge Stm., Kibweza Mt., 1720-1900 m, B&M 1680, Laula R., and rav., 1230 m, B&M 922, ibid. 1350-1400 m, B&M 1257, ibid. 1450-1550 m, B&M 1292, near Laula R. (BlZZARRI 1985), Musendemwa Stm., 1550-1850 m, B&M 1801; O 6: Kabinda Stm., Pweto - Kapulo 28 km, 960 m, B&M 1400; P5: Kasia R., Kieluzi -Mwela 40 km, 1780 m, B&M 1445, Kibobwa Mt. (BlZZARRI 1985), Lunangwa - Kab-wisi confl., 1820 m, B&M 1543, Pepa, 2000 m (KORNA$ 1979, BlZZARRI 1985); Q 6: Kiansamba R., spr. area, Kaliepe Mt., 1050-1200 m, B&M 3926. Habitat. Mostly lithophytic, in wet sites. Riparian and ravine dense forests, near - and amongst - waterfalls, on river banks, in fringing forests (on Biano Plateau), occasionally in open forests, wooded savannas (mostly on rocky slopes) and on termite mounds in forest environment. Found also on calcareous rocks, copper outcrops and moist sands; in deep or in slight shade. Zambia - Kornas (1979): 30. Selaginella cathedrifolia Spring Distribution. Fig. 13. Very rare, region of Tanganyika Lake, 1 station. O 1: Kabimba at Lubandaye R., 1200 m (Demaret 1955). Habitat. Terrestrial, riparian forest near spring, on moist soil. Zambia - Kornas (1979): not noted. Selaginella dregei (C. Presl) Hieron. Distribution. Fig. 14. Quite common, but only in eastern part of Upper Katanga, in the regions of the River Luapula and south-eastern border of “Sa-kania Boot” with Mount Mupwelume, alt. 950-1820 m (after 18 records), 19 stations. 30 N 11: Luapula R., Kyalwe, B&M 766, ibid. Johnston Falls 950 m, B&M 936, Lis. ¡0112, (SCHELPE 1973, KornaS 1979, Bizzarri 1985); P15: between Kipushia et Na-mopala, 1220 m (KornaS 1979, BIZZARRI 1985), Lundi R., 1220 m, B&M 1917, Mwala, 1235 m, B&M 2232, Namopala, 1300 m (Schelpe 1973, Bizzarri 1985); P 16: Kakulawele Mt., between Pemba and Lungo, 1400 m, B&M 2521, Kangili Mt., 1450 m, B&M 2506, Kasamba R. near spr., 1250 m, B&M 2735, Kasonso Mt., 1350 m and 1550 m, B&M 2567, 2724, Kiswanabantu Mt., 1350 m, B&M 2710, Kongolo Mt., between Ifuna and Malashi, 1320 m, B&M 2334, Malashi Mt., 1400 m, B&M 2707, Mubwe Mt., 1400 m, B&M 2513, Mupepe Mt., 1360 m, B&M 2717, Mupwelume Mt., 1820 m, B&M 2692, Serenje, 1370 m (SCHELPE 1973), Tchitwichasonfu Mt., 1455 m, B&M 2496. Habitat. Terrestrial or lithophytic, on/or between the rocks, mostly on rocky slopes, sometimes near river banks and in waterfalls, then on the rocks above water level, found also in moist meadow “dembo” on lateritic crusts; in full sunlight. Zambia - Kornas (1979): 31. SelagineUa kraussiana (Kunze) A. Braun Distribution. Fig. 15. Very rare, Muhila Plateau, alt. 1250-1740 m, 3 stations. O 4: Kiseye Mt., 1740 m, B&M 759, Laula R„ 1250 m, B&M 720 b, ibid. 1350-1400 m, B&M 1259 b, (Bizzarri 1985). Habitat. Terrestrial, found on steep montane slope and in ravine dense forest. Zambia - Kornas (1979): not noted. SelagineUa mittenii Bak. Distribution. Fig. 16. Rare, Biano and Muhila Plateaux, Kalemie region near Lake Tanganyika, “Sakania Boot” between Mount Mupwelume and Upper Luapula River, alt. 1400-1500 m (after 2 records), 5 stations. H 10: 2 km from Katentania, 1520 m (SCHELPE 1973); 110: 25 km from Katentania (Bizzarri 1985); N 1: Luama R., Tugulu (Demaret 1955); P4: Kampilikwe Mt. (Bizzarri 1985); P 15: Kiangulube Mt., 1400 m, B&M 1903. Beyond northern border of Upper Katanga: Lake Tanganyika, on Ubwari Peninsula (Demaret 1955). Habitat. Lithophytic or terrestrial, plant of very small size, recorded from moist rocks; usually grows on river banks and slopes, in gullies, found also in small montane forests (on Ubwari Peninsula). Zambia - Kornas (1979): 33. SelagineUa molliceps Spring Note. Voucher specimen B&M 1503 in KRA has the following remark of J. Kornas: “pas de fausses nervures”. Distribution. Fig. 17. Very rare, Biano and Marungu Plateaux - from there species determinations are doubtful, alt. 1640-1655 m, 3 stations. 31 111: Fungurume - Nguba (BlZZARRI 1985); P 5: Lunanga R., 1640 m, B&M 1503, Munamazi R„ env. Kanipini, 1655 m, B&M 1523 - on both recent stations S. cfr. mol-liceps. Habitat. No ecological information of Bizzarri l.c. Specimens of Selaginella found by B&M were growing in narrow fringing forest, limited by steep scarps. Zambia - Kornas (1979): not noted. Selaginella njam-njamensis Hieron. Distribution. Fig. 18. Very rare, Biano Plateau, 2 stations at the same altitude. 110: Mulamba Falls, 1450 m, B&M 685, Mulandakipindi Stm., 1450 m, B&M 698. Habitat. Litophytic, found on the rocks, in full sunlight. Zambia - Kornas (1979): 33. Selaginella shabaensis Bizzarri Note. Reported by Bizzarri (1982) as a new species, endemic to Upper Katanga. Distribution. Fig. 19. Very rare, west shore of Lake Tanganyika, in “locus classicus” only, 1 station. P3: Mpala (Bizzarri 1982, 1985). Habitat. Habitat not known. Zambia - Kornas (1979): not noted. Selaginella subisophylla Jermy Note. Endemic of Zambezi - Congo watershed (Kornas 1993 a). Determination of species according J. Kornas in KRA; the same collection number in BR was identified by Bizzarri (1985) as S. voluhilis Alston. Distribution. Fig. 20. Very rare, northern part of Kibara Plateau, 1 station. K 6: Mitwaba - Manono 30 km, Schm. 2041. Habitat. Terrestrial, found in seepage area among rocks. Zambia - Kornas (1979): 34. Selaginella tenerrima A. Braun ex Kuhn Distribution. Fig. 21. Very common, regions of rivers Lualaba and Upper Lufira, Biano and Muhila Plateaux, regions of Lake Moero, Lubumbashi and Mount Mupwelume - from there most numerous data, alt. 785-1500 m (after 37 records), 39 stations. G 10: Mutendele R., Kolwezi - Luena 114 km, 785 m, B&M 2042; I 10: Kadiva Falls, upstream of Kansenia, 1300 in, B&M 603; I 11: Fungurume “Hill V”, 1190 m, B&M 2063, Kimisombo Stm., Kakanda - Kambove 9 km, 1430 m, B&M 881; J 11: 32 Katuntukwa Rav., Kakanda - Kambove 13 km, 1440 m, B&M 874; J 12: Shinkolobwe, 1420 m, B&M 810; K 12: Katanga, 1140 m, B&M 906, Lupembashi R., env. Mangom-bo, 1140 m, B&M 915, 3 km ENE from Sofumwango railway station, Lubumbashi - Likasi 58 km, 1310 m, B&M 854; K 13: Fipango Stm., 1230 m, B&M 1112, Kafubu R., Kisanga, 1235 m, B&M 793, Kafubu R., cascades, 1230 m, B&M 774, Kansaba Stm., Kaponda, 1235 m, B&M 804, Kasapa R., Staquet Farm, 1220 m, B&M 2203, Lubumbashi (Bizzarri 1985), near Luiswishi Spr., Kiswishi Monastery, 1275 m, B&M 870; L 12: Luiswishi (Bizzarri 1985); L 14: Lubumbashi - Kasumbalesa, 1220 m, B&M 506; M 8: Luandashi R., Kyankalamu - Pweto 13 km, 985 m, B&M 1360; N 7: Mulonde R., Kyankalamu - Pweto 48 km, 1030 m, B&M 1384; O 4: Kabwa R., 1450 m, B&M 1230, Kansimba R„ 1425-1525 m, B&M 1155, Katwizia R„ 1400-1475 m, B&M 1182, Kibola R„ 1400-1475 m, B&M 1210, Laula Rav., 1350-1400 m, B&M 1254, Mutambala Stm., 1500 m, B&M 1158; P16: Akangili R., 1400 m, B&M 2438, Akasansa R. near spr., 1440 m, B&M 2554, Akeshimbi R., 1300 m, B&M 2375, Fibulukutu R., near spr., 1450 m, B&M 2606, Kamakolongo R., 1440 m, B&M 2489, Kasamba R., 1250 m, B&M 2741, Lungo R., 1370 m, B&M 2535, Mabondo R., between Ifuna and Malashi, 1290 m, B&M 2325, Malashi R., 1275 m, B&M 2345, Mpemba R., 1440 m, B&M 2474, Mubwe R., 1350 m, B&M 2423, Nyamanda R., 1290 m, B&M 2400, Tchitwichasonfu R., 1440 m, B&M 2457. Habitat. Terrestrial or lithophytic, in riparian and ravine dense forests - on river banks, rocks, rocky and sandy scarps, in narrow fringing forests and in open forests, found also on termite mounds, on brick wall of artificial dam and on mineralized outcrops at Shinkolobwe (Malaisse et al. 1994); mostly in shade. Zambia - Kornas (1979): 34. Selaginella versicolor Spring Distribution. Fig. 22. Rare, region of Lualaba River and north-eastern part of Upper Katanga: near Luvua River and Kalemie at Lake Tanganyika, on Mu-hila Plateau, alt. 785-1425 m (after 5 records), 8 stations. G 10: Mupwiji R., 800 m, B&M 2176, Mushinji R., Kolwezi - Kyamasumba 70 km, 1010 m, B&M 2157, Mutendele R., 810 m, B&M 365, ibid., Kolwezi - Luena 114 km, 785 m, B&M 2040; N 5: Mukui trib., 1290-1360 m, B&M 3628; O 1: Kavala Island, Golf of Bracone, Tanganyika Lake (Demaret 1955, BIZZARRI 1985); O 4: Nkala R., 1425 m, B&M 1351; O 5: Pweto - Moba road, between Katanti and Kalanga (Bizzarri 1985). Habitat. Terrestrial or epiphytic at tree bases: in riparian and ravine dense forests, occasionally on steep slopes and sandy scarps. Zambia - Kornas (1979): not noted. Selaginella sp. indet. Distribution. Specimens collected on Kibara and Muhila Plateaux and on southern periphery of Kundelungu Plateau, alt. 950-1250 m (after 3 records), 3 stations. 33 K 6: Mulumbi Esc., Mitwaba - Manono 52 km, 950 m, B&M 482\ L 11: Kaku R., 2 km N from Gombela, 1130 m, B&M 223\ O 4: Laula R., 1250 m, B&M 720 a, c. Habitat. Found in riparian forest, in ravines and in open forest - there on the steep, rocky slope. Equisetaceae Equisetum ramosissimum Desf. Distribution. Fig. 23*. Very rare, mostly in the region of Lubumbashi, found also near Kalemie at Lake Tanganyika, alt. 1150-1220 m (after 4 records), 7 stations. E. ramosissimum s. 1. K 13: Kisanga 1200 m (Schelpe 1973), Lubumbashi, near Gecamines Bridge (and mine), 1220 m, B&M 26, Lis. 10098, (Schelpe 1973); L 12: Kifumanzi R., Mututa, 1150 m (Schelpe 1973); O 1: Kalemie (Demaret 1955). E. ramosissimum subsp. ramosissimum Note. Taxonomic unit distinguished by some authors, among others J. Kornas. K 13: Kisanga (Lawalree 1969 b), env. Lubumbashi, 1200 m (Kornas 1979); O 1: Kalemie (Lawalree 1969 b) - other voucher specimen than quoted above after Demaret (1955) as E. ramosissimum s. 1. Habitat. Terrestrial, on river banks, scarps at the roads and disturbed sites under influence of industrial copper and cobalt pollution (cfr. Brooks et al. 1985), also in very moist meadows. Zambia - Kornas (1979): 35 - subsp. ramosissimum. Aspleniaceae Asplenium abyssinicum Fee Distribution. Fig. 24. Very rare, at Muhila Plateau, 1 station. O 5: Laula R„ 1250-1370 m, B&M 728. Habitat. Terrestrial, found in narrow ravine. Zambia - Kornas (1979): 94. Asplenium aethiopicum (Burm. f.) Becherer Distribution. Fig. 25. Very common, almost in the whole Upper Katanga, alt. 590-2020 m (after 134 records), 144 stations. G 11: Potopoto trib., 30 km SW from Kolwezi, 1425 m, B&M 339~, H 10: Katenta-nia, 1500 m (KORNAS 1979); H 11: Kolwezi - Mamfwe 12 km, 1350 m, B&M 2034\ 34 I 9: Dipera Falls, env. Lubudi, 1250-1310, B&M 427, 930, Kayo Falls, 1250-1310 m, B&M 418 bis; I 10: Gule R., 1450 m, B&M 536, Kalule Sud R„ 1500 m, B&M 613, Kipao R., 1590 m, B&M 652, Kipao Spr., 1630 m, B&M 554, Menge R., env. Quaglia Farm, 1575 m, B&M 572, Mulamba R., 1470 m, B&M 689, Paku trib., env. Biano Hotel, 1600 m, B&M 625; 111: Luita, 1500 m, B&M 892, Tshilongo Stm., 1300 m, B&M 328; J 11: 4-5 km N from Kakontwe, 1200 m, Mai. 10216, 10056, 4 km S from Kakontwe Mai. 9968, 6 km SE from Kambove, Kakontwe Rav., 1360 m, B&M 835; K 6: Kyowe R., Mitwaba - Manono 48 km, 1140 m, B&M 491, Manda R., Mitwaba - Manono 40 km, 1450 m, B&M 495; K 7: Bana Bankasi Hill, Mitwaba - Manono 11 km, 1600 m, B&M 459; K 10: Mwadingusha, 6 km from Lubumbashi - Likasi road bifurcation, 1325 m, B&M 397; K 12: Kamianga Stm., 5 km N from Shilatembo, 1200 m, B&M 284 a, Luembe R., near spr., 1275 m, B&M 54, Milondo Stm., 1190 m, B&M 2012 a, Mwera R., 1150 m, B&M 1128; K 13: Kisanga, 1200 m (Schelpe 1973), 11 km W from Lubumbashi, 1120 m (KORNAS 1979); L8: Kibwe wa Sanga Stm., Kimitondo, 1730 m, B&M 1039, Kwamba R„ 1650 m, B&M 1048, Mwanza R„ 1640 m, B&M 1029; L 9: Vibyo R., 1670 m, B&M 1070; L 10: Kapala R., 2 km N from Luishi, 1645 m, B&M 1017, 6 km N from Katshupa, 1690 m (SCHELPE 1973), Luanza Spr., 1690 m, B&M 984, Luanza Basin (KornaS 1979), Luanza R., Katshupa, 1680 m (Schelpe 1973), Lutsipuka Spr., 20 km SW from Msipashi, 1670 m (Schelpe 1973), Malando R., 1675 m, B&M 1004, Nungwe trib., 1670 m, B&M 1080; Petite Lofoi, 1550 m (KORNAS 1979); L 10?: Bembia Spr., W from Mwenge (region of middle Lufira R.), 1080 m (SCHELPE 1973); L 11: Kalembe R., 1550-1600 m (KORNAS 1979), Katwe R., spr. area, 1645 m, B&M 1105, Lutshipuka Falls, 1440-1550 m, B&M 232, 238, Mai. 11389, Mai. & Robbr. 1972, Lutshipuka Spr., 1670 m, B&M 967, Masansa Fall, 1500 m, B&M 276, Minwe Spr., 1620 m, B&M 962; L 12: Luiswishi, 1208 m, B&M 35, 204, 300, 516, Luiswishi R., 30 km NE from Lubumbashi, 1200 m, B&M 182; L 13: Lualala R., Lu-pembe, 1020 m (Schelpe 1973); L 14: env. Kasumbalesa, Lis. 10014, (KORNAS 1979); M 4: Kasenga Stm., 590 m, B&M 3175; M 5: Kakonkolwa Stm., 1190 m, B&M 3466, Kamabale Stm., 1200 m, B&M 3494, Konkwa Stm., 690 m, B&M 3242, Lukongolwe Stm., 1000 m, B&M 3430, Mbopofu R., 1250 m, B&M 3345, Mbula R., 700 m, B&M 3235, Mukalabafulue Stm., 900 m, B&M 3280, Tumengele R., 900 m, B&M 3410, ibid. A. cfr. aethiopicum B&M 3410 a; M 6: Lupwashi R„ 980 m, B&M 4174; M 7: Busa R. swamps, 960 m, B&M 4163, Kangidi R., 1000 m, B&M 4183; Kazamba R., 990-1250 m, B&M 4442, Mukolowanga R„ 990-1100 m, B&M 4491; M 8: Luandashi R., 985 m, B&M 1364; M 10: 6 km W from Kabiashia, 1350 m (Schelpe 1973), near Msipashi Fall, env. Kinika, 1250 m, Lis. 10002, (KORNAS 1979); N 4: Nungu R., 1300 m, B&M 3772; N 5: Lualashi R„ 1350-1410 m, B&M 3548, Mukui trib., 1285-1360 m, B&M 3631, 3649, 3688, 3700, Kalumbwe R„ B&M 1971, Kapundu, B&M 1979, Munshi R., B&M 1990; N 7: Kamipanga R., 975-1050 m, B&M 4532, Kasanza R„ 950-1050 m, B&M 4628, Kimapuya R„ 1250 m, B&M 4421, Mbikwa R., 975-1050 m, B&M 4567, Mateshi R„ 100-1150 m, B&M 4734, Mibiluka R„ 970-1050 m, B&M 4552; N 11: Johnston Falls, 950 m (Kornas 1979); N 15: Kalumbwe R., 1230 m, B&M 1971, Kapundu, 1230 m, B&M 1979, Munshi R„ 1225 m, B&M 1990; O 4: Ka-muinga R., 1725 m, B&M 1666, Kapondo R., 1500 m, B&M 1169, Kasenge Spr., Kib-weza Mt., 1720-1900 m, B&M 1695, Kilumbi R„ Kilenge Rap., 1400-1500 m, B&M 1316, 1837, 1842, Kipanda R„ 1550-1575 m, B&M 1200, 1204, Kisenga R., 1425 m, B&M 1865, Kitampa Spr., 1350-1400 m, B&M 1612, Kitu R., 1500 m, B&M 1747, Kizi 35 R., 1725 m, B&M 1671, Laula Rav., 1350-1400 m, B&M 1261, Luilu R., 1470 m, B&M 1797, Luilu trib., 1720-1820 m, B&M 1719, Lukonagulwa Stm., 1460 m, B&M 2825, Mazala R., 1700 m, B&M 1734, Miemba Spr., 1720-1750 m, B&M 1749, Mpa Stm., 1420 m, B&M 2940; O 4?: stm. near Kinianta, 1740 m, B&M 761; O 15: Kiangulube Stm., 1400 m, B&M 1907; P 4: Kije R„ 750-1130 m, B&M 3832, Kisambila Stm., 750-1130 m, B&M 3822; P 5: Kafupe Rav., 2000 m (KORNAS 1979), Kasia R., 1780 m, B&M 1447, Kazungalele R., 1500 m, B&M 4106, Kinsindwa R„ 2 km W from Kieluzi, 1750 m, B&M 1432, Kioma R„ 1380 m, B&M 4094, Lufuko R„ 1790-1840 m, B&M 1475, Lunanga R„ 1640 m, B&M 1512, Lunangwa - Kabwisi confl., 1820 m, B&M 1544, Lunkole Rav., Kieluzi - Mwela 45 km, 1890 m, B&M 1466, Luonde, 1900 m (Kornas 1979), Misongo Mt., 2020 m, B&M 3870; P 15: Kabeleshi R., 1230 m, B&M 2222, Kishitu Mt., 1240 m, B&M 1940, Lundi R., 1220 m, B&M 1918, Malebwe R., 1190 m, B&M 2236, Malembula Mt., 1240-1250 m, B&M 2214, Michinka Mt., 1280 m, B&M 1946, Mushampanga Mt., 1290 m, B&M 1933; P16: Akapemfia R., spr. area, Mupwelume Mt., 1500 m, B&M 2620, Akasansa R., spr. area, 1440 m, B&M 2558, Akeshimbi R., 1300 m, B&M 2382, Bulanga R., 1300 m, B&M 2360, Bulanga Mt., 1380 m, B&M 2697, Chinika R., spr. area, 1500 m, B&M 2657, Fibulukutu R., spr. area, 1450 m, B&M 2597, Ifuna R., Kindalo, 1240 m, B&M 2300, Kasala R„ 1245 m, B&M 2263 bis, Kasamba R., 1250 m, B&M 2746, Kasonso Mt., 1350 m, B&M 2722, Kimbwi R., spr. area, 1440 m, B&M 2576, Kiswanabantu Mt., 1350 m, B&M 2709, Kongolo Mt., between Ifuna and Malashi, 1320 m, B&M 2335, Lungo R., 1370 m, B&M 2537, Lusemfwa R., 1500 m, B&M 2639, Mabondo R., between Ifuna and Malashi, 1290 m, B&M 2326, Malashi Mt., 1400 m, B&M 2708, Mupwelume Mt., 1820 m, B&M 2695, Nyamanda R„ spr. area, Mupwelume Mt., 1530 m, B&M 2678, Umusangashi R., Mu-pala, 1305 m, B&M 2284; Q 6: Mukonge R., env. Sunkutu, 1100 m, B&M 4027; R 6: Kisenga R„ 750-1090 m, B&M 3982. Habitat. Lithophytic, terrestrial and occasionally epiphytic, in riparian, fringing and swamp forests, in ravine dense forests, mainly on moist rocks, in rock crevices or among boulders, near springs, on river banks, on rocks in river rapids and cascades; in woodlands on steep slopes, in wooded savannas on moist sites and on termite mounds. Some localities on calcareous formations. In shade, at alt. 2020 m in full sunlight. Zambia - Kornas (1979): 94. Asplenium sp. indet. - from A. aethiopicum complex Note. Here classified specimens, with black scales, belong possibly - according to the remark of J. Kornas in KRA - to a new species which deserves to be studied in detail (cfr. also the taxonomic remarks about A. aethiopicum in publication about Zambia (Kornas 1979). Distribution. Muhila and Marungu Plateaux, 2 stations. O 4: Musendemwa R., spr. area, 1550-1850 m, B&M 1821; P5: Munamazi R., env. Kanipini, 1655 m, B&M 1528. Habitat. Mostly terrestrial, found in ravine, in dry riparian forest on scarp and rocks and in narrow fringing forest limited by steep walls. Zambia - Kornas (1979): not noted. 36 Asplenium atroviride Schelpe Distribution. Fig. 26. Very rare, regions of River Lubule and of Lake Моего, alt. 940-1250 m (after 3 records), 3 stations. M 7: Kamitaba R., 990-1240 m, B&M 4468, Kazamba R., 990-1250 m, B&M 4435; N 7: Kita R., 940-1050 m, B&M 4613. Habitat. Terrestrial. In ravine dense forests, mostly on rocky scarps. Zambia - Korn as (1979): not noted. Asplenium blastophorum Hieron. Distribution. Fig. 27. Very rare, regions of rivers Upper Lufira and Luvua, area between Luvua River and Muhila Plateau, alt. 1300-1500 m (after 4 records), 4 stations. J 11: Kanduba R., 6 km W from Kakontwe, 1330 m, B&M 828; N 4: Kitebe R., 1300-1350 m, B&M 3806, Luhu R., 1475-1500 m, B&M 1336; N 5: Lualashi R„ 1300-1350 m, B&M 3072. Habitat. Epiphytic or terrestrial, found in riparian forests on river bank, on scarp, near spring and in the forest of shallow ravine. Zambia - Kornas (1979): 96. Asplenium buettneri Hieron. Distribution. Fig. 28*. Very common, regions of rivers Lualaba, Lufira, Luvua (most frequent), Lubule and Luapula, near Lakes Моего and Tanganyika, plateaux: Biano, Kundelungu, Muhila and Marungu - with peripheries, regions of Lubumbashi, Sakania and Mupwelume Mountain, alt. 550-1820 m (after 54 records), 61 stations. A. buettneri s. 1. G 10: Kyamasumba, 1080-1100 m, B&M 358, Mai. & Robbr. 2282; 110: Kansenia near Pande Dam, 1400 m, B&M 137, Kipao R., 1530 m, B&M 667; I 11: Fungurume - Nguba 9 km, 1220 m, B&M 2141, near Tshilongo Station, 1500 m, B&M 535; I 12: Swambo, B&M 1143; J 11: 8 km W from Kakontwe, 1330 m, B&M 828; К 9: Lufira Falls, Kiubo, 850 m (Schelpe 1973), Luvilombo R., Kiubo, 880 m, B&M 76; К 12: Milondo Stm., 3 km SW from Katanga, 1190 m, B&M 2012, 2019; К 13: Munama R., 10 km S from Lubumbashi (SCHELPE 1973), 22-28 km N from Lubumbashi, near the Farm Saint-Hubert, 1200 m, Lis. 10081, (SCHELPE 1973, KORNAS 1979); L4: Djipumba R., 550 m, B&M 3132; L 11: Katwe R., spr. area, 1645 m, B&M 1105; L 14: 20 km NNE from Tshinsenda, 1260 m (SCHELPE 1973); M 4: Djania R., 550 m, B&M 3125, Kalukenda Stm., 650 m - A. cfr. buettneri, B&M 3113, Kamahasa Stm., 590 m, B&M 3169, Katoma Stm., 610-620 m, B&M 3157, Kiminu R., 580 m, B&M 3194, Lusindoi R., 650 m, B&M 3098, Luvua R., 590 m, B&M 3147, Musinoi R., 590 m, B&M 3200; M 5: Imakabanga Stm., 690 m, B&M 3214, Kamianga - Lubalashi confl., 720-800 m, B&M 3448 - A. cfr. buettneri, Kamimvu Stm., 690 m, B&M 3208, Kyezimpala Stm., 690 m, B&M 3223, Luela R., 690 m, B&M 3251, Lukongolwe Stm., 1000 m, B&M 37 3430 a, Mbopofu R., 1250 m, B&M 3345 a, Mukalabafulue Stm., 900 m, B&M 3280 a- A. cfr. buettneri, Mukungai R., 690 m, B&M 3230; M 7: Lusalala R., Kyankalamu - Pweto 28 km, 1000 m, B&M 1369, 1370; N 4: Kamihafu R., 1320-1350 m, B&M 3726 - A. cfr. buettneri, Luiboso R., 1300 m, B&M 3778; N 5: Kapanitono, 1250 m, B&M 56, Mukui R., 1300-1360 m, B&M 3578; N 7: Loanza R„ 975-1050 m, B&M 4506, Lukonzolwa (KornaS 1979), Mulonde R., Kyankalamu - Pweto 48 km, 1030 m, B&M 1376; N 11: Johnston Falls, 950 m, B&M 938, 941; N 15: Sakania, 1250 m (Schelpe 1973); O 1: Kavala Island, Bracone Golf (Demaret 1955); O 2: Koki R., Mulange (Demaret 1955); O 4: Kalonga R„ 1390-1420 m, B&M 2884, Kapa Stm., 1720-1820 m,B&M 2791, KatomiaR., 1400 m, B&M 2964, KisengaR., 1425 m, B&M 1865, Laula R„ 1350-1400 m, B&M 1265, Luilu trib., 1720-1820 m, B&M 1719, Musendemwa Spr. and Stm. 1550-1850 m, B&M 1803, Muzimbwa R„ Makala Mt., 1650-1800 m, B&M 1648, Nkala R., 1425 m, B&M 1355; O 14: env. Kabunda, Giraud Falls (KORNAS 1979); O 15: Nkumbwa, 1230 m, B&M 1956; P 4: Kapupo Fall, Mulo-bozi R„ Kikarata, 1020-1100 m, B&M 2770, Kawa R„ 1280-1450 m, B&M 3846 b; P 5: 15 km from Kasiki (SCHELPE 1973), Malonge R., 1800 m, B&M4136; P6: Kapeta R., 1230 m, B&M 4068, Kapeta - Kibili confl., 1100 m, B&M 4078; P 16: Mpemba R., 1440 m, B&M 2475. Beyond Upper Katanga, near of its border: Vuano on Ubwari Peninsula, dry rocky hill (Demaret 1955). A. buettneri var. hildebrandtii Hieron. O 2: Koki R., Mulange in riparian forest (Demaret 1955) - near Lake Tanganyika. Beyond Upper Katanga: Vuano, Ubwari Peninsula, rocky hill (Demaret 1955). Habitat. Terrestrial or lithophytic, sometimes epiphytic. In riparian and ravine dense forests, mostly on rocky river banks and on steep rocky walls - found also in montane ravine forest with Juniperus procera, in dry dense forests “muhulu” and open forests on rocky slopes, on cupriferous hills and termite mounds; in shade and in open sites. Zambia - Kornas (1979): 97. Asplenium sp. indet. - ex aff. A. buettneri Hieron. (species nova or A. buettneri subsp.?) Note. According to some remarks of J. Kornas mentioned here specimens from KRA are without gemmae. Their determination is provisory and needs critical revision. Distribution. Region of Upper Lufira River, plateaux Kundelungu and Mu-hila - with periphery, alt. 1300-1620 m (after 4 records), 4 stations. K 12: Kabukande Stm., 2 km from the road to Sofumwango Station, 1300 m, B&M 861;\A\-. MinweSpr., 1620 m, B&M 962 bis;N4: Luhu R., 1475-1500 m, B&M 1336; O 4: Kibola R„ 1400-1475 m, B&M 1212. Habitat. Mostly epiphytic, found in ravine dense forest and in riparian and swamp forests. Zambia - KornaS (1979): 97 (A. buettneri). 38 Asplenium ceii Pic. Ser. Distribution. Fig. 29. Very rare, Muhila Plateau and between this area and Luvua River, alt. 1250-1450 m (after 4 records), 4 stations. N 5: Mukui trib., 1290-1360 m, B&M 3632; О 4: Laula Rav., 1250-1370 m, B&M 729, ibid. 1350-1400 m, B&M 1246, ibid. 1450 m, B&M 1284. Habitat. Epiphytic or lithophytic, in dense, ravine forets. Zambia - Kornas (1979): not noted. Asplenium chaseanum Schelpe Note. Endemic of Zambezi - Congo watershed (Kornas 1993 a). Distribution. Fig. 30. Quite common, regions of rivers Lualaba, Lufira, Lu-bule and Luapula, of Lake Моего and Lubumbashi, alt. 800-1325 m (after 17 records), 17 stations. G 10: Mupwiji R., 800 m, B&M 2173, Mushinji R„ Kolwezi - Kyamasumba 70 km, 1010 m, B&M 2169; К 10: Kikuswa Stm., 6 km from Mwadingusha, 1325 m, B&M 397; К 12: Kamianga Stm., 5 km N from Shilatembo, 1200 m, B&M 284; Luafi Stm., Lubumbashi - Likasi 70 km, 1230 m, B&M 306, Milondo Stm., 3 km SW from Katanga, 1190 m, B&M 2013, Mwera R„ 1150 m, B&M 2193, Shilatembo - to NW 5 km, 1340 m, Mai. & Robbr. 1731, Lukutwe Stm., 3 km ENE from Sofumwango Station, 1310 m, B&M 843; M 7: Kamitaba R„ 990-1240 m, B&M 4459, Kazamba R„ 990-1250 m, B&M 4445; N 7: Kamipanga R„ 975-1050 m, B&M 4526, Kitumbwe R., 1050 m (Schelpe 1973), Loanza R., 975-1050 m, B&M 4502, Mbikwa R., 975-1050 m, B&M 4568, Mibiluko R„ 970-1050 m, B&M 4549; N 10: Luapula R„ B&M 924. Habitat. Terrestrial or lithophytic, in riparian and in ravine dense forests, on rocky river borders, on scarps and - mostly - near temporary streams. Zambia - Kornas (1979): 97. Asplenium cfr. christii Hieron. Note. Determination of J. Kornas in KRA is preliminary, with the following remark about studied voucher specimen: “basal sterile leaves lacking, leaf apices broken, no gemmae visible, more material needed!”. Distribution. Fig. 31. Very rare, Muhila Plateau, 1 station. О 4: Laula R„ 1370 m, B&M 734. Habitat. Terrestrial, found in narrow ravine. Zambia - Kornas (1979): not noted. Asplenium dregeanum Kunze Distribution. Fig. 32. Quite common, but only in the north-eastern part of Upper Katanga, on plateaux: Muhila with its western periphery and Marungu, alt. 1230-1450 (1900) m (after 12 records), 12 stations. 39 N 4: Kajia R., 1300-1350 m, B&M 3008, Kamihafu R., 1320-1350 m, B&M 3743, Kitebe R., 1300-1350 m, B&M 3812, Luiboso R., 1300 m, B&M 3791, Nungu R., 1300 m, B&M 3764-, N 5: Kibondo R„ 1335 m, B&M 3034, Lualashi R„ 1300-1350 m, B&M 3069, Mukui R„ 1300-1360 m, B&M 3599, Mukui trib., 1285-1360 m, B&M 3633, 3662, 3683, 3707', О 4: Laula R„ rav., 1230-1250 m, B&M 786 bis, 727, ibid. 1350-1450 m, B&M 730, 731b, 731c, 732,1255, 1276\ P 5: Kibobwa Rav., foots of Kibobwa Mt., 1900 m, Lis. Mai. & Sym. 11932. Habitat. Terrestrial or epiphytic, in riparian and in ravine dense forests, on rocks and rocky walls, at river banks and occasionally at waterfalls. Zambia - KornaS (1979): 98. Asplenium elliottii С. H. Wright Distribution. Fig. 33. Rare, Biano, Kundelungu and Marungu Plateaux, alt. 1470-1890 m (after 9 records), 9 stations. 110: Kalule Sud R„ 1500 m, B&M 619, ibid. 1565 m (SCHELPE 1973), ibid. Biano Hotel, 1530 m (SCHELPE 1973), Mulamba R., 1470 m, B&M 688\ L 10: Nungwe trib., 1670 m, B&M 1074\ P 5: Lufuko R., 7 km from Pepa, 1790-1840 m, B&M 1470, Lun-kole Rav., Kieluzi - Mwela 45 km, 1890 m, B&M 1456, Malonge R., 1800 m, B&M 4136, Mufufu R„ 1880 m, B&M 1565. Habitat. Mostly epiphitic (in forest undergrowth), in fringing, riparian and swamp forests, near river rapids and waterfalls, on scarps, on very moist sands, in shade. Zambia - Kornas (1979): not noted. Asplenium erectum Bory ex Willd. Distribution. Fig. 34. Quite common, Kundelungu Plateau, region of Luvua River, Muhila and Marungu Plateaux - most numerous records, ait. 1270-1940 m (after 12 records), 12 stations. L 10: Malando R., near spr., 7 km NNE from Katshupa, 1690 m (SCHELPE 1973); N 5: Mukui trib., 1285-1360 m, B&M 3623, 3650, 3684, 3709', О 4: Kalusembe R„ 1270-1380 m, B&M 2834, Kapa Stm., 1720-1820 m, B&M 2786, Kapondo R„ 1500 m, B&M 1180 a-A. cfr. erectum, Libanda R., 1450 m, B&M 2905, Luilu trib., 1720-1820 m, B&M 1716, Lukongolwe R., 1280-1360 m, B&M 2856', P 5: Lufuko R., 1790-1940 m, B&M 1473, Lunanga R., 1640 m, B&M 1506, Lunkole Rav., Kieluzi - Mwela 45 km, 1890 m, B&M 1458, Munamazi R„ 1655 m, B&M 1520. Habitat. Terrestrial, occasionally epiphytic, in riparian and ravine usually dense forests, in fringing forests, on steep scarps, rocky walls, near springs, river rapids and waterfalls. Zambia - Kornas (1979): 98 - var. usambarense. Asplenium formosum Willd. Distribution. Fig. 35. Very common, Biano, Kundelungu and Muhila Plateaux, regions of Lufira River, Lubumbashi, of Lake Моего and Luvua River; 40 most numerous quotations from north-eastern part of Upper Katanga, alt. 680-1850 m (after 64 records), 72 stations. G 11: Nzilo Rav. (DE Wildeman 1927, after SCHELPE 1973); H 11: Kolwezi - Mamf-we 12 km, 1350 m, B&M 2032; 19: Dipera Falls, 1220 m, B&M 424, Kayo Falls, 1250— 1310 m, B&M 417, Kalule Nord, Kayo, 1390 m, B&M 2, Lubudi - Nguba 27 km, 1400 m, B&M 442; 110: Gule R., 1450 m, B&M 544, Kalule Sud, 10 km from Lubudi, 1270 m (SCHELPE 1973), Kipao R„ 1530 m, B&M 659, Mulamba R., 1470 m, B&M 694, Pande Fall, upstream of Kansenia, 1300 m, B&M 592; 111: Luita trib., 1480 m, B&M 902; J 11: Kanduba R., 8 km W from Kakontwe, 1330 m, B&M 824, Likasi (Kornas 1979); К 10: Kikuswa Stm., 6 km from Mwadingusha, B&M 403; К 12: Luafi Stm., rav., Lubumbashi - Likasi 70 km, 1230 m, B&M 305, Milondo Stm., 3 km SW from Katanga, 1190 m, B&M 2008, Mwera R., 1150 m, B&M 2194; К 13: Nkashima R., near Kilando, 1260 m (SCHELPE 1973), Kisanga R., Kisanga, 1200 m (SCHELPE 1973); L 10: Lofoi Vai., upstream of Kaloba Fall, 1200 m, B&M 163, Munyankara Stm., env. Gom-bela, 1060 m, B&M 394; L 11: Kaku R., 2 km N from Gombela, 1130 m, B&M 228, Masansa Fall, 1500 m, B&M 275; M 5: Kakonkolwa Mt. and Stm. Mt., 1190 m, B&M 3471, Kamabale Stm., 1200 m, B&M 3481, Kamisamba R., 900 m, B&M 3307, Kima-tambo Stm., 1350 m, B&M 3395, Lubalashi - Luvua confl., 680 m, B&M 3259, Lukon-golwe Stm., 1000 m, B&M 3428, Mbopofu R., 1250 m, B&M 3342, 3351, Muhezie R., 1000 m, B&M 3367, Mukalabafulue Stm., 900 m, B&M 3282, Tumengele R., 900 m, B&M 3411; M 10: Luanza R., 4 km WNW from Kabiashia, 1130 m (SCHELPE 1973), env. Kinika, 1180m (KORNAS 1979); N 4: Kajia R„ 1300-1350 m, B&M 3018, Kamihafu R., 1320-1350 m, B&M 3745, Kitebe R„ 1300-1350 m, B&M 3808, Luiboso R„ 1300 m, B&M 3794, Luhu R„ 1475-1500 m, B&M 1331, Nungu R„ 1300 m, B&M 3770; N 5: Kasenga R„ 1300-1340 m, B&M 3560, Kibondo R., 1335 m, B&M 3033, Lualashi R„ 1300-1350 m, B&M 3071, ibid, near spr., 1350-1410 m, B&M 3545, Mukui R., 1300-1360 m, B&M 3579, Mukui trib., 1290-1360 m, B&M 3618, 3651, 3686, 3712; N 7: Mulonde R., Kyankalamu - Pweto 48 km, 1030 m, B&M 1375; О 1: Коку Val., 20 km from Kalemie (SCHELPE 1973), Tugulu (Demaret 1955); О 2: Koki R., Mulange (Demaret 1955); О 4: Kabwa R„ 1450 m, B&M 1227, Kalonga R„ rav., 1390-1420 m, B&M 2885, Kalusembe R„ 1270-1380 m, B&M 2833, Кара Stm., 1720-1820 m, B&M 2790, Kapondo R„ 1500 m, B&M 1180, Katomia R„ 1400 m, B&M 2966, Kilumbi R„ Kilenge Rap., 1450-1500 m, B&M 1305, Kisenga R., 1425 m, B&M 1867, Kitampa Stm., spr. area, 1350-1400 m, B&M 1620, Laula R., 1250 m, B&M 724,1256, Libanda R., 1450 m, B&M 2904, Lukongolwe R., 1280-1360 m, B&M 2855, Miemba Stm., spr. area, 1720-1750 m, B&M 1752, Mpa Stm., 1420 m, B&M 2934, Musendemwa Stm., 1550-1850 m, B&M 1805, 1820, near Mutungulu R., 1430 m, B&M 738, Muzimbwa R„ 1650-1800, B&M 1641, Nkoko Stm., 1350 m, B&M 2989; P 4: Mulobozi R„ Kapupo Fall, 1020-1100 m, B&M 2779. Beyond the northern border of Upper Katanga: Vuano, near Ubwari Peninsula (Demaret 1955). Habitat. Lithophytic or epiphytic, in riparian (most often) and ravine dense forests, near waterfalls, on river banks, along streams (seasonal and perrenial), on rocky scarps and rocks, occasionally in fringing forests and in open forests on rocky slopes. Zambia - Kornas (1979): 99. 41 Asplenium friesiorum C. Chr. Distribution. Fig. 36. Rare, mostly Kundelungu Plateau, one quotation from Marungu Plateau, alt. 1650-1880 m (after 9 records), 10 stations. L 8: Kwamba R„ 1650 m, B&M 1044; L 9: Vibyo R„ 1670 m, B&M 1069; L 10: Luanza Basin, Katshupa (Malaisse 1975, KornaS 1979), Luanza Spr., 1680-1690 m, B&M 250, 989, Luanza R., 1655 m, B&M J001, ibid. 6 km N from “Poste de Katshupa”, 1660 m (SCHELPE 1973), 20 km SW from Msipashi, 1675 m (Schelpe 1973), Nungwe trib., 1670 m, B&M 1072-, L 11: Lutshipuka Spr., 1670 m, B&M 970; P 5: Mufufu R„ 1880 m, B&M 1562. Habitat. Terrestrial, in riparian and swamp forests, near springs, in very moist sites, occasionally in narrow fringing forests. Zambia - Kornas (1979): 99. Asplenium inaequilaterale Willd. Distribution. Fig. 37. Very common (in some parts of Upper Katanga): northern periphery of Biano Plateau, regions of Lubumbashi and of Luvua River, Muhila Plateau - most numerous data, and Marungu Plateau, alt. 680-1890 m (after 33 records), 33 stations. I 9: Kalule Nord R„ Kayo Falls, 1250-1310 m, B&M 921, ibid. 1340 m, B&M 3; K 12: Luafi Stm., Lubumbashi - Likasi 70 km, 1230 m, B&M 317; M 5: Kamabale R., 1350-1420 m, B&M 1627, Kimatambo Stm., 1350 m, B&M 3387, Lubalashi - Luvua confl., 680 m, B&M 3263, Lukongolwe Stm., 1000 m, B&M 3429, Mbopofu R., 1250 m, B&M 3349, Muhezie R., 1000 m, B&M 3370, Mukalabafulue Stm., 900 m, B&M 3279, Musanikwa R„ 800-1000 m, B&M 3327, Tumengele R., 900 m, B&M 3406; N 4: Kamihafu R., 1320-1350 m, B&M 3744, Luiboso R., 1300 m, B&M 3789, 3792, Nungu R„ 1300 m, B&M 3771; N 5: Kasenga R„ 1300-1340 m, B&M 3561, Lualashi R„ 1350-1410 m, B&M 3546, Mukui trib., 1300-1360 m, B&M 3652, 3713, Mukui R., 1300-1360 m, B&M 3585, ibid. B&M 3582 - A. cfr. inaequilaterale; O 4: Kalusembe R., 1270-1380 m, B&M 2834, Kansimba R., 1425-1525 m, B&M 1149, Kapondo R„ 1500 m, B&M 1164, Kisenga R., 1425 m, B&M 1864, Laula R., rav., 1350-1400 m, B&M 1250, Libanda R„ 1450 m, B&M 2905, Lukongolwe R., 1280-1360 m, B&M 2856, Mpa Stm., 1420 m, B&M 2935; P4: Mulobozi R., Kapupo Fall, 1020-1100 m, B&M 2772 a; P 5: Lunkole Stm., Kieluzi - Mwela 45 km, 1890 m, B&M 1457, Mumbulwa R„ 1750 m, B&M 4117; Q 6: Kiansamba R„ Kaliepe Mt., 1050-1200 m, B&M 3940, Mukonge R„ Kibwebwe Fall, 1190-1300 m, B&M 4054, Mu-konge R., 1100 m, B&M 4031. Habitat. Terrestrial or occasionally epiphytic, in riparian (most often) and ravine dense forests, on steep rocky sites as e.g. river banks, near waterfalls and springs. Zambia - Kornas (1979): 100. Asplenium lademannianum Rosenst. Distribution. Fig. 38. Rare, Biano and Kundelungu Plateaux, Muhila Plateau and region of Luvua River, alt. 1220-1670 m (after 6 records), 6 stations. 42 I 9: Dipera Falls, 1220 m, B&M 429, Kayo Falls, 1250-1310 m, B&M 418; L 9: VibyoR., 1670 m, B&M 1070 a; NS: Mukui R., 1300-1360 m, B&M 3578 bis - 4. cfr. lademannianum, Mukui trib., 1290-1360 m, B&M 3631 a; O 4: Kabwa R., 1450 m, B&M 1231. Habitat. Epiphytic or lithophytic, in riparian and ravine dense forests, on rocky walls, rocks in the rivers and near waterfalls. Zambia - Kornas (1979): not noted. Asplenium lunulatum Sw. Distribution. Very rare, known in two varieties from the banks of Lake Tanganyika. A. lunulatum var. erectum (Bory) Sim (= A. erectum Bory ex Wilid.) Note. Beyond the area of present study: Ubwari Peninsula (Demaret 1955). Zambia - Kornas (1979): 98 (Asplenium erectum var. usambarense). A. lunulatum var. minor Sim Distribution. Fig. 39. Very rare, 1 station. N 1: Tendele R., Tugulu (Demaret 1955). Habitat. On steep banks of the river. Zambia - Kornas (1979): not noted. Asplenium mannii Hook. Distribution. Fig. 40. Very rare, Kundelungu and Muhila Plateaux, alt. 1320-1670 m (after 4 records), 4 stations. L 9: Vibyo R., 1670 m, B&M 1063', N 4: Kamihafu R., 1320-1350 m, B&M 3749, Luhu R„ 1475-1500 m, B&M 1335; O 4: Libanda R„ 1450 m, B&M 2923. Habitat. Mostly epiphytic, in riparian forests. Zambia - Kornas (1979): 100. Asplenium monanthes L. Distribution. Fig. 41. Very rare, region of Muhila and Marungu Plateaux, alt. (1320) 1655-1820 m (after 3 records), 3 stations (one with doubtful determination of species). N 4: Kamihafu R., 1320-1350 m, B&M 3746 - A. cfr. monanthes; P 5: Lunangwa - Kabwisi confl., 1820 m, B&M 1542, Munamazi R., env. Kanipini, 1655 m, B&M 1517. Habitat. Mostly terrestrial, in narrow fringing forests; specimen with uncertain determination was found at relatively low altitude, on the rocks in the riparian forest. Kornas (1979): 101 - in Zambia not noted, quoted from the adjacent areas in Malawi and Rhodesia (Zimbabwe). 43 Asplenium normale D. Don. Note. Distinguished by J. Kornas in some preliminary way. In publication of Johns (1991) this taxon is included in “a list of the species as recorded in Peter (1929)”, with a remark that many specific names listed in Peter’s paper do not correspond to those applied at present in Africa. Distribution. Fig. 42. Very rare, region of Luvua River and western periphery of Muhila Plateau, 2 stations at the same altitude. N 4: Kitebe R„ 1300-1350 m, B&M 3807, 3810; N 5: Lualashi R., 1300-1350 m, B&M 3073. Habitat. Lithophytic? In riparian forests, on river banks and rocky scarps. Zambia - Kornas (1979): not noted. Asplenium paucijungum Ballard Distribution. Fig. 43. Very rare, announced from 1 station. O 1: Lumandaye R., Kabimba, 1200 m (DEMARET 1955). Habitat. Terrestrial, on moist sites in riparian forest. Zambia - Kornas (1979): not noted. Asplenium preussii Hieron. (= A. pseudoauriculatum Schelpe) Taxonomic remark. A. preussii is only slightly distinct from A. pseudoauriculatum and in some recent publications, e.g. Burrows (1990), names of both species are considered as synonyms. Distribution. Fig. 44. Very rare, Muhila Plateau, alt. 1370-1450 m, 2 stations. O 4: Laula R., rav. 1370 m, B&M 735, ibid, on the level of the waterfall, 1400-1450 m, B&M 1279. Habitat. Terrestrial, found in the ravine, on sandy and rocky soil. Zambia - Kornas (1979): not noted. Asplenium protensum Schrad. Distribution. Fig. 45. Common, but only in north-eastern part of Upper Katanga: Marungu - and mostly Muhila Plateaux, region of Luvua River, alt. 900-1900 m (after 27 records), 27 stations. M 5: Kamisamba R., 900 m, B&M 3303, Kimatambo Stm., 1350 m, B&M 3385, Lu-kongolwe Stm., 1000 m, B&M 3433, Mbopofu R., 1250 m, B&M 3350, Tumengele R., 900 m, B&M 3405; N4: Kamihafu R., 1320-1350 m, B&M 3725, Luhu R., 1475-1500 m, B&M 1323; N 5: Mukui R„ 1300-1360 m, B&M 3584, Mukui trib., 1290-1360 m, B&M 3615;O4; KamabaleR., 1350-1420 m, B&M 1631, Kamuinga R„ 1725 m, B&M 44 1661, Kapa Stm., 1720-1820m, B&M 2787, Kapondo R„ 1500 m, B&M 1168, Kasenge Stm., spr. area, 1720-1900 m, B&M 1684, Kilumbi R., Kilenge Rap., 1400 m, B&M 1844, Kipanda R., 1550-1575 m, B&M 1201 - A. cfr. protensum, Kiseye Mt., rav. near Kinianta, 1730 m, B&M 757, Kitampa Stm., spr. area, 1350-1400 m, B&M 1614, Laula R., rav., 1250 m, B&M 723, ibid. 1350-1400 m, B&M 1245, Libanda R., 1450 m, B&M 2903, Luilu R., 1470 m, B&M 1796, Luilu trib., 1720-1820 m, B&M 1713, Musendem-wa Stm., spr. area, 1550-1850 m, B&M 1806, 1822-, P 5: Luande, 1860 m (Kornas 1979), Lunanga R., 1640 m, B&M 1509, Lunkole Rav., Kieluzi - Mwela 45 km, 1890 m, B&M 1460. Habitat. Terrestrial, occasionally epiphytic or lithophytic, in riparian and in ravine dense forests, on river banks, rocks and steep slopes (sometimes on sandy walls), found also in fringing forest. Zambia - Kornas (1979): 101. Asplenium pumilum Sw. subsp. hymenophylloides (Fee) Schelpe Note. Taxon announced as new for Upper Katanga (Shaba) by Kornas et al. (1990). All stations mentioned below are quoted in that publication. Therefore citation is ommited in the following list, with exception of cases when other information (e.g. from KRA Herbarium) was not accessible. Distribution. Fig. 46. Common, Biano and Muhila Plateaux, southern periphery of Marungu Plateau, regions of rivers upper Lufira and Luapula, of Lu-bumbashi, Sakania and Mount Mupwelume, alt. 1140-1525 m (after 25 records), 25 stations. I 9: Dipera Falls, 1220 m, B&M 430, Kayo Falls, 1250-1310 m, B&M 415; I 10: Kadiva Falls, 1300 m, B&M 605, Pande Fall, 1300 m, B&M 589, 602-, 1 11: Kimisombo Stm., Kakanda - Kambove 9 km, 1430 m, B&M 880; J 11: 6 km SSE from Kambove, 1360 m, B&M 840; K 12: Kapandashiba R„ 3 km WNW from Katanga, 1150 m, B&M 2003, Luembe Riv., spr. area, 1275 m, B&M 51, Lukunka R., 1170 m, B&M 1124, Lu-pembashi R., env. Mangombo, 1140 m, B&M 917, Mwera R., 1170 m, B&M 2192; K 13: Kafubu R., cascades, 1230 m, B&M 779; M 14: Luina R., Kibomboma Falls, 1200 m (KORNAS et al. 1990); N 15: Kalumbwe R., 1230 m, B&M 1970, Kapundu, 1230 m, B&M 1982, Lukusashi R., 1230 m, B&M 1962, Munshi R„ 1225 m, B&M 1984; O 4: Kansimba R., 1425-1525 m, B&M 1154; O 15: Menda, 1230 m (Kornas et al. 1990), Nkumbwa, 1230 m, B&M 1958; P 16: Akangili R., 1350 m, B&M 2434, Kamakolongo R„ 1440 m (Kornas et al. 1990), Mubwe R., 1350 m, B&M 2412, Tchitwichasonfu R., 1140 m (KORNAS et al. 1990); Q 6: Kiansamba Spr., Kaliepe Mt., 1050-1200 m, B&M 3924. Habitat. Terrestrial, lithophytic or occasionally epiphytic, mostly in riparian forests, near waterfalls, on scarps and on the termite mounds situated in part in open woodlands, in fringing forests; found also in shallow gully on calcareous rocks (calcaire de Kakontwe) and on artificial wall of the bridge. Zambia - Kornas (1979): 102. 45 Asplenium sandersonii Hook. Distribution. Fig. 47. Very rare, Muhila Plateau, alt. 1350-1400 m, 2 stations. O 4: Laula R., rav., 1370 m, B&M 731a, ibid., 10 m from the waterfall, 1350— 1400 m, B&M 1251. Habitat. Terrestrial, found in narrow ravine, on rocky wall. Zambia - Kornas (1979): not noted. Asplenium cfr. schelpei Braithw. Note. Determination very doubtful, needs revision as stressed in KRA J. Kornas, who left also a remark: “not A. stuhlmanii (rhachis green & scaly)”. Taxonomic problems concerning this taxon are also mentioned by Kornas (1979) in reference to Zambia. Distribution. Fig. 48. Recorded from southern periphery of “Sakania Boot”, mainly from Mount Mupwelume, alt. 1190-1305 m (after 6 records), 6 stations. O 15: Kiangulube Stm., 1300 m, B&M 1894-, P 16: Kasala R., 1245 m, B&M 2263, Malashi R„ 1275 m, B&M 2352, Malebwe R., 1190 m, B&M 2250, Umusangashi R., Mupala, 1305 m, B&M 2284, Umwimbwa R., Mupala, 1300 m, B&M 2276. Habitat. Terrestrial or epiphytic, found in riparian forests at river banks, on rocky slope and between the rocks. Zambia - Kornas (1979): 103. Asplenium theciferum (Kunth) Mett. Note. Var. concinnum (Schrad.) Schelpe, distinguished by some authors (and by J. Kornas in KRA), is mentioned below separately. According to former publication (Kornas 1979) it is however a “taxon in need of critical revision”. Distribution. Fig. 49*. Very common, regions of rivers Lualaba, Lufira and Luvua, plateaux: Biano, Kundelungu, northern periphery of Kibara, Muhila - most numerous data - and Marungu, Plain of Lubumbashi, alt. 900-1900 m (after 40 records), 61 stations. A. theciferum s. 1. G 10: Kyamasumba, 1080 m, B&M 357, Kyamasumba - Kolwezi 13 km, 1075 m, B&M 2143, 7 km from Luilu Stm., near Kolwezi, 1250 m, B&M 353; G 11: Luilu Vai. near Kolwezi, 1160 m, B&M 346; I 9: Dipera Falls, 1220 m, B&M 425, Kayo Falls, 1250-1310 m, B&M 409; I 10: Gule R„ 1450 m, B&M 540, Kalule Sud R., 1500 m, B&M 623, ibid, near Biano Hotel, B&M 641, (Schelpe 1973 - as var. concinnum), Kansenia, near Pande Dam, B&M 138, Kipao Spr., 1630 m, B&M 552, Kipao R., 1530 m, B&M 658, Mulamba R., 1470 m, B&M 682; K 10: 6 km from Mwadingusha, B&M 402; L 6: Luiza R„ Kisele - Mitwaba road, 1530 m, B&M 90; L 8: Kwamba R„ 1650 m, B&M 1047; L 9: Vibyo R., 1670 m, B&M 1066; L 10: Nungwe trib., B&M 1081; L 11: Katwe R., B&M 1104, Lutshipuka Spr., 1670 m, B&M 968, Lutshipuka 46 Falls, 1520 m, B&M 240, Minwe Spr., 1620 m, B&M 957; L 12: Luiswishi, 1208 m, B&M 517; M 5: Mukalabafulue Stm., 900 m, B&M 3293; N 4: Luiboso R., 1300 m, B&M 3790, Luhu R„ 1475-1500 m, B&M 1333; N 5: Mukui trib., 1285-1360 m, B&M 3664, 3689, 3714; O 4: Kabwa R„ 1450 m, B&M 1234, Kalusembe R„ 1270-1380 m, B&M 2849, Kapondo R., 1500 m, B&M 1178, Kasenge Spr., Kibweza Mt., 1720-1900 m B&M 1694, Katomia R„ 1400 m, B&M 2980, Kibola R„ 1400-1475 m, B&M 1217, Kilumbi R„ Kilenge Rap., 1400-1500 m, B&M 1307, 1858, Kipanda R„ 1550-1575 m, B&M 1200, Kisenga R., 1425 m, B&M 1881, Kitampa Stm., spr. area, 1350-1400 m, B&M 1621, Kizi R„ 1725 m, B&M 1670, 1673, Laula Rav., 1350-1400 m, B&M 1259, Libanda R., 1450 m, B&M 2924, Lukonagulwa Stm., 1460 m, B&M 2829, Lukongolwe R., 1280-1360 m, B&M 2868, Mpa Stm., 1420 m, B&M 2951, Musen-demwa Stm., 1550-1850 m, B&M 1830, Muzimbwa R., 1650-1800 m, B&M 1643, Nkala R„ 1425 m, B&M 1356, Nkoko Stm., 1350 m, B&M2999; PS: Lufuko R„ 1790-1840 m, B&M 1479, Lunkole Stm., 1890 m, B&M 1468, Mufufu R„ 1880 m, B&M 1558. A. theciferum var. concinnum (Schrad.) Schelpe G 11: Kolwezi, 1100 m (KORNAS 1979), Nzilo Rav. (DE WlLDEMAN 1923, DE Wil-DEMAN, Staner 1932, after SCHELPE 1973); 111: Dipeta R„ Fungurume, 1210-1220 m (Schelpe 1973, Kornas 1979); K 13: KisangaR., near Lubumbashi, 1200 m (Schelpe 1973), 11 km from Lubumbashi, 1220 m (KORNAS 1979), 20 km from Lubumbashi, 1250 m (Schelpe 1973); L10: 6 km N from Katshupa (Schelpe 1973), near Lofoi Spr., 1600 m, Lis. 10062, (Kornas 1979), Petite Lofoi R., 1550 m, Lis. Mai. & Sym. 12525 a, (Kornas 1979), Luanza catchment area (Malaisse 1975, Kornas 1979); L 13: Lua-lala R., Lupembe, 1020 m (SCHELPE 1973) Habitat. Epiphytic or lithophytic, in riparian, swamp and ravine dense forests, occasionally in fringing forests and dense dry forests “muhulu”, mostly on river banks, scarps and near seasonal streams. Zambia - Kornas (1979): 103 - A. theciferum var. concinnum. Asplenium unilaterale Lam. Distribution. Fig. 50. Rare, near Lake Tanganyika, on Muhila Plateau, in the region of Lubule River and Lake Moero, alt. 990-1820 m (after 7 records), 8 stations. M 7: Kamitaba R., 990-1240 m, B&M 4471; N 7: Bupompo R„ 1000-1100 m, B&M 4667, Kimapuya R„ 1250 m, B&M 4426; O 1: M’Toa (Demaret 1955); O 4: Kapa Stm., 1720-1820 m, B&M 2794, Laula R„ rav., 1370 m, B&M 736, ibid. 1450-1550 m, B&M 1285, Luilu trib., 1720-1820 m, B&M 1710. Habitat. Epiphytic or lithophytic. In wooded savanna (near Lake Tanganyika), in ravine dense forests on scarps and occasionally on emerged rocks among the river; in shade. Zambia - Kornas (1979): not noted. Asplenium sp. indet. - 3 groups of specimens with mentioned below remarks of J. Kornas: 1) “ A. sp., not A. stuhlmanii - rhachis green and scaly” 47 Distribution. Found in the region of Sakania and - mostly - of Mupwe-lume Moutain, alt. 1190-1305 m, 6 stations. O 15: Kiangulube Stm., 1300 m, B&M 1894; P 15: Malebwe R., 1190 m, B&M 2250; P 16: Kasala R„ 1245 m, B&M 2263, Malashi R„ 1275 m, B&M 2352, Umusan-gashi R., Mupala, 1305 m, B&M 2284, Umwimbwa R., Mupala, 1300 m, B&M 2276. Habitat. These ferns were growing in riparian forests as epiphyte, or between the rocks and on rocky slopes at river banks. 2) “A. sp.: damaged leaves and one gemma” - 1 specimen, and “without gemmae” - 9 specimens. Distribution. Found in the region of Luvua River and Muhila Plateau, alt. 590-1820 m, 10 stations. M 4: Mangendwe Stm., 590 m, B&M 3184; M 5: Lubalashi - Luvua confl., 680 m, B&M 3265, Muhezie R., 1000 m, B&M 3369, Musanikwa R„ 800-1000 m, B&M 3326; N 5: Kibondo R„ 1335 m, B&M 3037; O 4: Kalusembe R„ 1270-1380 m, B&M 2840, Kapa Stm., 1720-1820, B&M 2793, Nkoko Stm., 1300 m, B&M 2992, Libanda R„ 1450 m, B&M 2906, Lukongolwe R., 1280-1360 m, B&M 2863. Habitat. These fems were growing in riparian and occasionally in ravine dense forests, mostly on river baks and scarps. 3) A. sp. (without any remark) and *A. sp. iuv. Distribution and Habitat. Periphery of Marungu Plateau, Biano Plateau. I 9: Dipera Falls, env. Lubudi, 1220 m, B&M 933* at the base of tree trunks and on dry rocks; N 5: Mukui R., 1300-1360, B&M 3598 on the scarp in riparian forest. Azollaceae Azolla nilotica Decne. ex Mett. Distribution. Fig. 51. Very rare, region of Lufira River, 2 stations. K 9 ?: Lufira R. (LAWALREE 1976); K 9: Lufira R., Kalanga, terr. Mitwaba (Lawalree 1976). Habitat. Aquatic, floating in shallow water, occasionally amphiphytic. Zambia - Kornas (1979): 122. Azolla pinnata R. Br. var. africana (Desv.) Bak. (= A. africana Desv.) Distribution. Fig. 52. Quite common, Kibara and Marungu Plateaux, regions of rivers Lufira, Luvua, Lubule and Luapula and of Lake Moero, Plain of Lubumbashi, alt. 900-1800 m (after 4 records), 16 stations. J 11: Kapolowe, Lake Tshangalele (Lawalree 1976); K 9: Lufira R., Kiubo (Lawalree 1976), ibid., Kiubo - Shimana (Schelpe 1973, Lawalree 1976); K 13: Kafubu R., 48 near Kasungami, B&M 501, (SCHELPE 1973), Karavia R., B&M 515, Kimilolo R., B&M 129, Kisanga, env. Lubumbashi, Lis. 10113, (Lawalree 1976, KORNAS 1979), Lubum-bashi (SCHELPE 1973, Lawalree 1976); M 4: Kamahasa Stm., B&M 3165, Luvua R., B&M 3142; M 7: Kangidi R„ 1000 m, B&M 4186; N 6: Pweto, 900 m (SCHELPE 1973, Lawalree 1976); N 10: Kasenga (Lawalree 1976); N 11: Johnston Falls, Luapula (Lawalree 1976), Mibuyu R., 950 m, B&M 945; P 5: Bonanza Swamp, 1800 m, Lis. Mai. & Sym. 10232, (KORNAS 1979). Habitat. Aquatic - floating on shallow water in quiet bays of lakes and rivers (also in riparian forests), in swamps and in flooded meadows. Zambia - Kornas (1979): 123. Blechnaceae Blechnum australe L. var. australe Note. Species announced as new for Upper Katanga (Shaba) by Kornas et al. (1990). Station mentioned below is quoted in that publication. Distribution. Fig. 53. Very rare, Marungu Plateau, 1 station. P5: Munamazi R., env. Kanipini, 1655 m, B&M 1516. Habitat. Terrestrial, found in narrow fringing forest limited by steep walls. Zambia - Kornas (1979): not noted. Blechnum tabulare (Thunb.) Kuhn Distribution. Fig. 54. Common, plateaux: Manika, Biano, Kundelungu (most numerous data), Kibara and Marungu, region of Mount Mupwelume, alt. 1370-2250 m (after 22 records), 26 stations. G 11: Manika Plateau (Lawalree 1971, Kornas 1979); H 10: Katentania R., 2 km from “Poste de Katentania”, 1500 m, B&M 715; I 10: Biano II, 1560 m, B&M'630, Kipao R., 1590 m, B&M 648; К 7: Bana Bankasi Hill, Mitwaba - Manono 11 km, 1600 m, B&M 458, Mitwaba - Manono 9 km, 1500 m, B&M 498; L 8: Lufukwe R., 1650 m, B&M 1059; L9: Kamakanga R., Luizi, 1600 m (SCHELPE 1973); L 10: Kapala R., 2 km N from Luishi, 1645 m, B&M 1015, Luanza R., 6 km N from Katshupa 1680 m, B&M 259, ibid. Luanza trib. 1670 m (SCHELPE 1973), Luanza Basin, Katshupa (Lawalree 1971, Malaisse 1975, Kornas 1979), Luanza R., spr. area, 1690 m, B&M 978; L 11: Kalembe R., 1550 m (KORNAS 1979), Katwe R. (LAWALREE 1971, KORNAS 1979), Katwe Spr., 16 km SW from Msipashi (SCHELPE 1973); PS: Kasiki, 2250 m (Kornas 1979), Kioma R., 50 km SE from Pepa, 1380 m, B&M 4095; P16: Akapemfia R., spr. area, Mupwelume Mt., 1500 m, B&M 2625, Akasansa R., spr. area, 1440 m, B&M 2555, Chinika R., spr. area, Mupwelume Mt., 1500 m, B&M 2666, Fibulukutu R., spr. area, 1450 m, B&M 2603, Kimbwi R., spr. area, Mupwelume Mt., 1440 m, B&M 2581, Lungo R., 1370 m, B&M 2540, Lusemfwa R., 1500 m, B&M 2645, Nyamanda R., spr. area, Mupwelume Mt., 1530 m, B&M 2683. 49 Habitat. Terrestrial, on wet sites, in swamp forests near springs, in fringing and mostly in riparian forests, on river banks, scarps, on the rocks or between them, sometimes on sand. Noted also in wooded savanna, on seepage area and on the slope covered by rocky blocks. Zambia - Kornas (1979): 119. Cyatheaceae Cyathea dregei Kunze Distribution. Fig. 55. Very common, regions of rivers Lualaba, Luvua and of Lake Moero, plateaux: Biano, Kibara, Kundelungu, Muhila and Marungu, regions of Lubumbashi and of Mount Mupwelume, alt. 1000 - 2020 m (after 54 records), 57 stations. G 11: 23 km NNW from Kolwezi, 1350 m, Mai. & Robb. 2200; H 10: Katentania R., 2 km from “Poste de Katentania”, 1500 m, B&M 716; I 10: Kipao R., 1590 m, B&M 644, Kisamoya R., 2 km ESE from Biano Railway Station, 1600 m (SCHELPE 1973), Menge R., env. Quaglia Farm, 1575 m, B&M 566, 576, Mulamba Falls, 1450 m, B&M 670, 4 km from Mulamba Falls, 1700 m, B&M 130, Paku trib. env. Biano Hotel, 1600 m, B&M 624; K 6: Lusinga R., Lusinga 1780 m (Schelpe 1973), Mitwaba - Ma-nono 12 km, 1630 m, B&M 460; K 12: Luafi Stm., Lubumbashi - Likasi 70 km, 1230 m, B&M 313; K 13: 15 km SE from Lubumbashi, 1150 m (SCHELPE 1973); L 8: Kimitondo Stm., 1730 m, B&M 1036, Lufukwe R„ 1650 m, B&M 1060; L 10: Kapala R„ 2 km N from Luishi, 1645 m, B&M 1013, Katshupa, 1560-1620 m (Malaisse 1975, Kornas 1979), ibid., 6 km N from “Poste de Katshupa”, Luanza R„ val„ 1670 m, B&M 256 b, 260, Mai. 5235, (SCHELPE 1973), Luanza R., 1655 m, B&M 998, Luanza Basin (Malaisse 1975, Kornas 1979), 12 km WNW from Msipashi, 1700 m (SCHELPE 1973), Nungwe trib., 1670 m, B&M 1089; L 11: 15 km SW from Msipashi, Katwe R., 1660 m (SCHELPE 1973); L 13: Munama R., Kasamba, 1220 m, B&M 1581; M 5: Lukongolwe Stm., 1000 m, B&M 3437; N 5: Kaha R„ 1290-1320 m, B&M 3534, Kilungu Stm., 1075-1100 m, B&M 3513, Lualashi R„ 1350-1410 m, B&M 3541, Mukui trib., 1285-1360 m, B&M 3680, 3702; N 7: Mutemuna R., Kyankalamu - Pweto 77 km, 1290 m, B&M 1386; O 4: Kamakubi R., 1500 m, B&M 1742, Kamuinga R., 1725 m, B&M 1664, Kansimba R., spr. area, 1465 m, B&M 1779, Kapa Stm., 1720-1820 m, B&M 2801, Katomia R„ 1400 m, B&M 2970, Katwizia R„ 1400-1475 m, B&M 1188, Kilumbi R., Kilenge Rap., 1400 m, B&M 1852, Kitampa Stm., 1350-1400 m, B&M 1617, Kipanda R„ 1550-1575 m, B&M 1205, Kisenga R„ 1425 m, B&M 1871, Kizi R„ 1725 m, B&M 1667, Lukonagulwa Stm., 1460 m, B&M 2822, Mwango Mt., 1770 m, B&M 762, Nkala R., 1425 m, B&M 1349, Nkuntwi R., Milembwa, at Pweto - Muhila road, Mai. 11947; O 6: Kafibukulo Stm., Pweto - Musosa 13 km, 1690 m, B&M 1417; P4: Liapen-da Rav., 45 km from Moba (SCHELPE 1973); P 5: Kitendwe R„ Kampazi, 1815 m (Schelpe 1973), Lunanga R., Kamipini, 1650 m (SCHELPE 1973), Lunanga R„ 1640 m, B&M 1504, Lunangwa - Kabwisi confl., 1820 m, B&M 1546, Misongo Mt., 2020 m, B&M 3875; P 16: Akapemfia R„ spr. area, 1500 m, B&M 2622, Akasansa R., spr. area, 1440 m, B&M 2561, Fibulukutu R., spr. area, 1450 m, B&M 2605, Kimbwi R., spr. area, 1440 m, B&M 2583, Lungo R., 1370 m, B&M 2543, Lusemfwa R., 1500 m, B&M 2647. 50 Habitat. Tree fern, in riparian (most often), fringing and ravine dense forests, near waterfalls, on river and stream banks, on swampy sites near springs, occasionally on slopes; on rocky or sandy soils, in shade, exceptionally in open sites. Zambia - Kornas (1979): 80. Cyathea thomsonii Bak. Distribution. Fig. 56. Quite common, mostly in north-eastern part of Upper Katanga: on Muhila Plateau with western periphery and near Luvua River, found also on Kundelungu Plateau, alt. (680) 1270-1820 m (after 12 records), 13 stations. L 10: Luanza Basin (Malaisse 1975, Kornas 1979), Luanza R., 6 km N from “Poste de Katshupa”, 1680 m (Malaisse 1969, Schelpe 1973); M 5: Lubalashi -Luvua confl., 680 m, B&M 3268; N 4: Kajia R., 1300-1350 m, B&M 3012; N 5: Ki-bondo R., 1335 m, B&M 3036, Lualashi R., 1300-1350 m, B&M 3068; O 4: Kalonga Rap., 1390-1400 m, B&M 2880, Kalusembe R., 1270-1380 m, B&M 2845, Luilu trib., 1720-1820 m, B&M 1706, Libanda R„ 1450 m, B&M 2912, Lukongolwe R„ 1280-1360 m, B&M 2858, Nkoko Stm., 1350 m, B&M 2995; O 6: Kyamba Rav., road crossing Kapulo to Pweto - Musosa 9 km, 1650 m, B&M 1409. Habitat. Tree fern, in riparian and occasionally in ravine dense forests, mostly on river banks, near river rapids and springs, on moist slopes and in swampy places. Zambia - Kornas (1979): 81. Davalliaceae Davallia chaerophylloides (Poir.) Steud. Distribution. Fig. 57. Very rare, region of Lualaba River, 2 stations; 2 other stations beyond the border of Upper Katanga. G 10: Kolwezi - Kyamasumba 70 km (Lawalree 1993), near Wakipinji, Kolwezi - Kyamasumba 112 km, 850 m, B&M 2148, (Lawalree 1993). Beyond Upper Katanga - E 5: Ruashi R., Kaniama, B&M 925, Kasese, B&M 188; J 5: Ngoy Mani, at the road Kabumbulu - Kabongo, 670 m (SCHELPE 1973). Habitat. Epiphytic, found in open forest - near Wakipinji and in cluster of Platycerium elephantotis - in riparian forest at Ruashi River. Zambia - Kornas (1979): not noted. Dennstaedtiaceae Blotiella crenata (Alston) Schelpe Distribution. Fig. 58. Common, mostly plateaux: Biano, Kundelungu, Muhila - with some periphery, also Manika, Kibara and Marungu and region of the River Luapula, alt. 1300-1880 m (after 29 records), 30 stations. 51 G 11: Munzenzi R., Muanamoya, 1480 m, Lis. Mai. & Sym. 4198; 110: Kalule Sud trib., Biano Hotel, 1560 m (SCHELPE 1973), Kipao R., spr. area, 1630 m, B&M 551, Menge R., env. Quaglia Farm, 1575 m, B&M 573, road of Quaglia Farm, 1670 m, B&M 141; K 6: Kanda Stm., Mitwaba - Manono 34 km, 1400 m, B&M 463; L 8: Kimitondo Stm., Kibwe wa Sanga, 1730 m, B&M 1041, Kwamba R., 1650 m, B&M 1042, 1056; L10: Luanza Basin, Katshupa (Malaisse 1975, KornaS 1979), Luanza R., 6 km NNW from Katshupa 1690 m (SCHELPE 1973), Luanza Spr., 1690 m, B&M 980, Malando R., spr. area, 1675 m, B&M 1008, 20 km SW from Msipashi, 1675 m (SCHELPE 1973), Nungwe trib., 1670 m, B&M 1082; L 11: Luanza R., val., 6 km N from Katshupa, 1680 m, B&M 251, Lutshipuka Spr., 1670 m, B&M 977, Minwe Spr., 1620 m, B&M 960; M 12: 8 km from Kiniama, 1620 m (SCHELPE 1973); N 4: Kajia R., 1300-1350 m, B&M 3015; O 4: env. Kansimba, 1380 m, B&M 737, Kapa Stm., rav., 1720-1820 m, B&M 2800, Katomia R., 1400 m, B&M 2971, Kipanda R„ 1550-1575 m, B&M 1197, Libanda R„ 1450 m, B&M 2907, Lukonagulwa Stm., 1460 m, B&M 2827, Mpa Stm., 1420 m, B&M 2946, Nkoko Stm., 1350 m, B&M 2996; O 6: Kyamba Rav., from road crossing Kapulo to Pweto - Musosa 9 km, 1650 m, B&M 1410; P 5: Mufufu R., 1880 m, B&M 1568; Q 6: Kinsamba R., Kaliepe Mt., 1050-1200 m, B&M 3937. Habitat. Terrestrial, on wet sites in riparian, fringing, and in dense forests growing on the slopes and in ravines, in swamp forests - near springs and river banks, on tree foots; occasionally on sandy soils and in forest clearings. Zambia - Kornas (1979): 85. Blotiella currorii (Hook.) Tryon Distribution. Fig. 59. Rare, plateaux Kundelungu and Muhila - with its western peripheries, alt. 1270-1670 m (after 7 records), 7 stations. L 10: Katwe R., spr. area, 1645 m, B&M 1102, Nungwe trib., 1670 m, B&M 1084; N 5: Kibondo R„ 1335 m, B&M 3043, Lualashi R„ 1300-1350 m, B&M 3060; O 4: Kalusembe R., 1270-1380 m, B&M 2836, Katomia R., 1400 m, B&M 2972, Libanda R„ 1450 m, B&M 2913. Habitat. Terrestrial, in riparian forests, mostly on river banks, also on swampy sites near springs. Zambia - Kornas (1979): 86. Blotiella glabra (Bory) Tryon Distribution. Fig. 60. Very rare, Marungu Plateau, 1 station. P5: 15 km from Kitendwe, 1900 m (SCHELPE 1973). Habitat. Terrestrial, found in dense thickets growing on swamp edges. Zambia - Kornas (1979): 86. Blotiella cfr. natalensis (Hook.) Tryon Note. Determination of species is uncertain, as marked J. Kornas in KRA. Distribution. Fig. 61. Very rare, 1 station, on Kundelungu Plateau. L 11: Katwe R., spr. area, 1645 m, B&M, 1101, 1102 bis. 52 Habitat. Terrestrial, found in riparian forest, swampy in places. Zambia - Kornas (1979): 86. Histiopteris incisa (Thunb.) J. Sm. Distribution. Fig. 62. Quite common, mostly on the plateaux: Manika, Biano, Kundelungu, Muhila and Marungu, alt. 1190-1880 m (after 19 records), 20 stations. G 11: Mwanamoya, 1480 m (KornaS 1979); I 10: Biano Hotel, 1570 m (SCHELPE 1973), Kalule Sud trib., 1575 m (SCHELPE 1973), Kitanalesa R„ 1570 m, B&M 700, Menge R., 1575 m, B&M 582, Mulamba Falls in a distance of 4 km, 1700 m, B&M 144; L 8: Kwamba R„ 1650 m, B&M 1055; L 10: 6 km N from Katshupa, 1690 m (SCHELPE 1973), Luanza Basin (Malaisse 1975, KORNAS 1979), Luanza R., val., 6 km N from Katshupa, 1680 m, B&M 252, Luanza Spr., 1690 m, B&M 983, Malando Spr., 1675 m, B&M 1003, 12 km WNW from Msipashi, 1700 m (SCHELPE 1973), Nungwe trib., 1670 m, B&M 1085; L 11: Katwe R., spr. area, 1645 m, B&M 1100, Lutshipuka Spr., 1670 m, B&M 971; O 4: Mpa Stm., 1420 m, B&M 2938; PS: Mufufu R„ 1880 m, B&M 1571; Q 6: Kiansamba R., spr. area, Kaliepe Mt., 1050-1200 m, B&M 3939, Mukonge R„ Kibwebwe Fall, env. Musali, 1190-1300 m, B&M 4058. Habitat. Terrestrial, in riparian and swamp - mostly montane - forests, near springs, found also on the rocks near waterfall; in wet sites and soils rich in humus. Zambia - KornaS (1979): 84. Hypolepis sparsisora (Schrad.) Kuhn Distribution. Fig. 63. Very rare, Marungu Plateau, 1 station. PS: Mufufu R„ 1880 m, B&M 1554. Habitat. Terrestrial, found in swamp forest at the beginning of exploitation. Zambia - Kornas (1979): not noted. Microlepia speluncae (L.) Moore Distribution. Fig. 64. Common, plateaux: Biano, Kundelungu and Muhila with its western periphery, periphery of Marungu Plateau, regions of Lakes Tanganyika and Moero, of Luvua River and Plain of Lubumbashi, alt. 650-1690 m (after 25 records), 26 stations. I 10: Menge R., near Quaglia Farm, 1575 m, B&M 578, 923; K 12: Luafi Stm., Lubumbashi - Likasi 70 km, 1230 m, B&M 307, Luembe R., near spr., 1275 m, B&M 47; K 13: Kafubu R., env. Lubumbashi, 1210 m, B&M 294, Kasungami, 1215 m, B&M 202;'L8; Kwamba R„ 1650 m, B&M I046;lAfr. Luanza Spr., 1690 m, B&M 986; L 11: Luanza R., val., 6 km N from Katshupa, 1680 m, B&M 254; L 12: Kibalashi R., Sapwe, 1090 m, B&M 1888; M 5: Kasa Mt. and Stm., 650-690 m, B&M 3082; N 4: Kijiya R., sector Muneke, 1400-1425 m, B&M 1344; N 5: Kibondo R„ 1335 m, B&M 3041, Mukui R., 1300-1360 m, B&M 3590; N 7: Mutemuna R., Sudi, Kyankalamu - Pweto 53 77 km, 1290 m, B&M 1392; O 4: Kalonga R., 1390-1420 m, B&M 2878, Kansimba R„ 1465 m, B&M 1782, KatomiaR., 1400 m, B&M 2969, KipandaR., 1550-1575 m, B&M 1203, Kitampa Stm., spr. area, 1350-1400 m, B&M 1611; Laula R., rav., 1350-1400 m, B&M 1271, Luhu R„ 1475-1500 m, B&M 1326 a, Lukongolwe R„ B&M 2861, Mpa Stm., 1420 m, B&M 2943, Nkoko Stm., 1350 m, B&M 2990; Q 6: Nsamba R„ spr. area, Kaliepe Mt., 1050-1200 m, B&M 3933; R 6: Kita R„ 920 m, B&M 4007. Habitat. Terrestrial, in riparian and less often in ravine dense forests, also in some disturbed forest stands, on river and stream banks, on sandy scarps, in swamp forests around springs, in swamps and edges of wet meadows “dembo”; in shade, occasionally in full sunlight. Zambia - KornaS (1979): 81. Odontosoria africana Ballard Distribution. Fig. 65. Very rare, region of Kalemie at Lake Tanganyika and Muhila Plateau, alt. 1200-1820 m (after 3 records), 3 stations. O 1: Lubandaye R., Kabimba, 1200 m (Ballard 1957, Kramer 1971); O 4: Kapa Stm., 1720-1820 m, B&M 2805, Luilu trib., 1720-1820 m, B&M 1702. Habitat. Terrestrial (climbing-according Johns 1991), found in riparian forest - at Lubandaye River, and in ravine forests on steep bank and moist scarp - on other localities. Zambia - KornaS (1979): not noted. Odontosoria chinensis (L.) J. Sm. var. divaricata Christ (= Stenoloma chinense (L.) Bedd. var. divaricatum (Christ) Alston) Distribution. Fig. 66. Very rare, near Lake Tanganyika, 1 station. O 1: Lubandaye R., Kabimba, 1200 m (Demaret 1955). Habitat. Found in riparian forest. Zambia - Kornas (1979): not noted. Pteridium aquilinum (L.) Kuhn Taxonomic remark. In publication of 1979 Kornas wrote: “the Zambian plants represent two morphologically distinct types, which do not integrate with one another”. In spite of this among voucher specimens from Upper Katanga he distinguished two subspecies and also the intermediate forms, quoted below. Some voucher specimens were however left without subspecies distinction. P. aquilinum s. 1. Distribution. Fig. 67*. Pteridium aquilinum is very common in Upper Katanga and was reported from 105 stations (taking into account intraspecific units). Data for specimens classified to species level only are the following: Kundelungu and Muhila Plateaux, region of Luapula River, alt. 1220-1670 m, 3 stations. 54 L 10: Nungwe trib, 1670 m, B&M 1076 (iuv.); L 13: Lwamisamba Stm., Sambwa Mission, 1220 m, B&M 1598; O 4: env. Kilenge Rap., 1400 m, B&M 1831. Beyond the border Upper Katanga: near Lake Tanganyika, on Ubwari Peninsula (Demaret 1955). Habitat. Terrestrial. Found near spring in riparian forest, swampy in places, on the edge of reed community and in pasture at the rock foots; on Ubwari among the savanna. Zambia - KornaS (1979): 82. P. aquilinum subsp. aquilinum Distribution. Fig. 68 (map includes also the intermediate form). Very common, in various parts of Upper Katanga: region of Lualaba River, plateaux: Biano, Kundelungu, Marungu, Muhila - and its periphery, regions of Luvua River, Lake Moero, Plain of Lubumbashi and south-eastern border of “Sakania Boot” with Mount Mupwelume, alt. 720-2060 m (after 49 records), 49 stations. G 10: Mutendele R., 810 m, B&M 366', G 11: Potopoto trib., 30 km SW from Kol-wezi, 1425 m, B&M 337; H 10: Katentania R., 1500 m, B&M 708; I 10: Biano Hotel, 1570 m (Schelpe 1973), 8 km from Kiniama, W from the road Biano - Lubudi, 1630 m (Schelpe 1973), Kitanalesa R., 4 km from Quaglia Farm, 1570 m, B&M 702, Menge R., env. Quaglia Farm, 1575 m, B&M 565; K 12: Sofumwango, 2 km from the road to the station, 1300 m, B&M 858; K 13: near Lubumbashi, 1220 m (SCHELPE 1973); L8: Kimitondo Stm., Kibwe wa Sanga, 1730 m, B&M 1037, Kwamba R., 1650 m, B&M 1054; L 9: Kamakanga R., Luizi 1600 m (SCHELPE 1973); L 10: Kapala R., 2 km from Luishi, 1645 m, B&M 1019, 6 km NNW from “Poste de Katshupa”, 1680 m (Schelpe 1973), Luanza Basin, Katshupa, 1690 m (MALAISSE 1975, KORNAS 1979); L 11: Luan-za Spr., 1690 m, B&M 982, Lutshipuka Spr., 1670 m, B&M 972, Minwe Spr., 1620 m, B&M 952; M 5: Kamabale Stm., 1200 m, B&M 3476, Kamianga - Lubalashi confl., 720-800 m, B&M 3441, Mukalabafulue Stm., 900 m, B&M 3273, Tumengele R., 900 m, B&M 3401; M 8: Luandashi R„ Kasolo, Kyankalamu - Pweto 13 km, 985 m, B&M 1359; N 4: Luhu R„ 1475-1500 m, B&M 1330; N 5: Kasenga R„ 1300-1340 m, B&M 3552, Mukui R„ 1300-1360 m, B&M 3568; O 4: Kamuinga R„ 1725 m, B&M 1652, Kapa Stm., 1720-1820 m, B&M 2782, Kibola R., 1400-1475 m, B&M 1211, Ki-lumbi R„ Kilenge Rap., 1450-1500 m, B&M 1298, Kipanda R„ 1550-1575 m, B&M 1195, Kitampa Stm., spr. area, 1350-1400 m, B&M 1608, Laula R„ rav., 1350-1400 m, B&M 1242, Luilu trib., 1720-1820 m, B&M 1712, Lukongolwe R„ 1280-1360 m, B&M 2851, Lukonagulwa Stm., 1460 m, B&M 2814, Mazala R„ 1700 m, B&M 1726, Mpa Stm., 1420 m, B&M 2930, Musendemwa Stm., 1550-1850 m, B&M 1810, Nkala R., 1425 m, B&M 1346; P 4: Kampilikwe, W from the road Moba - Pepa, 2000 m (SCHELPE 1973), S from Kampilikwe, E from the road Moba - Pepa, 2060 m (SCHELPE 1973), Mulobozi R„ Kapupo Fall, 1020-1100 m, B&M 2761, Kawa R„ 1280-1450 m, B&M 3842; P5: Kitendwe, SW from the road Moba - Pepa, 1830 m (SCHELPE 1973), Misongo Mt., 2020 m, B&M 3872, 5 km ESE from Pepa, 1990 m, B&M 3859; P 15: Malebwe R., 1190 m, B&M 2242; P 16: Akapemfia R., spr. area, 1500 m, B&M 2612. Habitat. Terrestrial, in dense and in open forests, mostly on edges of riparian forests, also in ravine and fringing forests, in wooded savannas and grass 55 lands (“steppes”), in disturbed forests, among thickets e.g. bordering swamp on lateritic crusts, occasionally on the rocks; in shade and in full sunlight. Zambia - Kornas (1979): 83 (subsp. aquilinum var. aquilinum). P. aquilinum subsp. aquilinum ad subsp. centrali-africanum Hieron. vergens Distribution. That intermediate form was found in the region of Lualaba River and on Muhila Plateau with its periphery, alt. 1250-1400 m, 3 stations. H 11: 12 km E from Kolwezi, 1400 m, B&M 772; N 5: Kapanitono, 1250 m, B&M 931; O 4: Lukongolwe R„ 1280-1360 m, B&M 2870. Habitat. Terrestrial. In open woodlands on rocky slopes (between the rocks). Observed also as epiphyte in some riparian forest. Zambia - Kornas (1979): form not noted. P. aquilinum subsp. centrali-africanum Hieron. (= P. aquilinum var. caudatum (Linné) Tardieu - name used by DEMARET 1955). Distribution. Fig. 69 (map includes also the intermediate form). Very common, distributed similarly as subsp. aquilinum, but found also on Kibara Plateau and near Sakania, alt. 950-1820 m (after 43 records), 48 stations. H 10: NW from Ditanto R., 5 km from Katentania, 1530 m (Schelpe 1973), Katen-tania R., 2 km from “Poste de Katentania”, 1500 m, B&M 710; I 10: 2 km from Kan-senia, Biano Road, 1550-1600 m (Schelpe 1973), Pande Vai. (Schelpe 1973); J 12: Shinkolobwe, 1420 m, B&M 806; K 7: Mitwaba, 1600 m, B&M 450; K 12: Katanga, 1140 m, B&M 912, Lukunka R., 1170 m, B&M 2189; K 13: Kanakolwe Stm., 1225 m, B&M 155, 156, Kisanga near Lubumbashi, 1200 m (Schelpe 1973), Lubumbashi R. (left), near Boitsfort Farm, 1210 m (SCHELPE 1973), 10 km from Lubumbashi, N from Kasapa Road, 1240 m (SCHELPE 1973), 12 km NW from Lubumbashi, 1300 m (Schelpe 1973), 15 km from Lubumbashi, E from the road to “La Chaumière” (Schelpe 1973), near Nkingandu Mission, 1225 m, B&M 30; L 10: Kapala R., about 2 km N from Luishi, 1645 m, B&M 1027, 6 km NNW from Katshupa, 1690 m (SCHELPE 1973), Luanza Basin (MALAISSE 1975, KORNAS 1979); L 11: Minwe Spr., 1620 m, B&M 956; L 12: Luiswishi R., 30 km NE from Lubumbashi, 1200 m, B&M 179; L 13: Luiswishi R., Mususwa, 1200 m, B&M 794, Munama R., Kasamba, 1220 m, B&M 1578; L 14: 4 km N from Kasumbalesa, 1500 m, B&M 93, Kinsenda, 1330 m, B&M 2084, Tshinsenda (Tshisinka) (DE Wildeman 1923, DE Wildeman, Staner 1932, after Schelpe 1973); N 5: Kabanga R„ 1175-1200 m, B&M 3517, Kaha R„ 1290-1320 m, B&M 3526, Kilungu Stm., 1075-1100 m, B&M 3507, Lualashi R„ 1300-1350 m, B&M 3051, ibid. 1350-1410 m, B&M 3536; N 15: Kalumbwe R„ 1230 m, B&M 1978, Mun-shi R., 1225 m, B&M 1989; O 4: Kalusembe R., 1270-1380 m, B&M 2831, Kansimba R., spr. area, 1465 m, B&M 1774, Kansimba trib., 1490 m, B&M 1762, Kapa Stm., 1720-1820 m, B&M 2781, Kilumbi R„ Kilenge Rap., 1450-1500 m, B&M 1300, Kyula Mt., 1700 m, B&M 752, Luilu R„ 1470 m, B&M 1790, Luilu trib., Lukanda Swamp, 1470 m, B&M 1786, Lukonagulwa Stm., 1460 m, B&M 2813, Musendemwa Stm., spr. area, 1550-1850 m, B&M 1799; O 6: Mukosi R., near Kibwe, 950 m, B&M 1402; P 4: 56 Mulobozi R., Kapupo Fall, 1020-1100 m, B&M 2760; P 15: Kabaleshi R., 1230 m, B&M 2224, Michinka Mt., 1280 m, B&M 1945, Mushampanga Mt., 1290 m, B&M 1932', P 16: Kimbwi R., spr. area, B&M 2570. Beyond Upper Katanga: near Lake Tanganyika, on Ubwari Peninsula, in savanna (Demaret 1955), and M 1: Kioko-Nyumba, in the riparian forest (Demaret 1955). Habitat. Similar sites that those of P. aquilinum subsp. aquilinum. Recorded mostly at forest edges, found also as synanthropic plant e.g. on gravel heaps. KornaS (1979): 83. P. aquilinum subsp. centrali-africanum ad subsp. aquilinum vergens Distribution. This intermediate form was found on Marungu Plateau, on 1 station. P 5: 5 km ESE from Pepa, 1990 m, B&M 3858. Habitat. Thickets at seasonal swamp occurring on lateritic crust. Zambia - KornaS (1979): form not noted. Dryopteridaceae Athyrium annae Kornas Note. Endemic of the Zambezi - Congo watershed (Kornas 1993 a), announced as new for Upper Katanga (Shaba) by Kornas et al. (1990). All stations mentioned below on the basis of KRA herbarium are quoted in that publication. Its citation is therefore ommited with two exceptions concerning some additional data. Distribution. Fig. 70. Rare, regions of Lualaba and Lufira Rivers, Plain of Lubumbashi, alt. 800-1310 m (after 7 records), 7 stations. G 10: 7 km from Luilu Vai. near Kolwezi, Nzilo - Kyamasumba 18 km, 1230-1250 m, B&M 351, Mupwiji R., 800 m (KornaS et al. 1990); H 10: Lutungulu R., Kasofu - Sokele 7 km, 890 m (KORNAS et al. 1990); K 10: Shilatembo - Mwadingusha 6 km, 1300 m, B&M 400; K 12: Luafi Stm., Lubumbashi - Likasi 70 km, 1230 m, B&M 315, 2023, Mai. 10852, ibid, near Mangombo, 1300 m, Mai. & Robbr. 2132, 3 km ENE from Sofumwango Station, 1310 m, B&M 845. Habitat. Terrestrial or lithophytic. Mostly in ravine dense forests, found also on the boulders near seasonal stream. Zambia - Kornas (1979): 104. Athyrium scandicinum (Willd.) C. Presl. Distribution. Fig. 71. Rare, plateaux: Kundelungu and Muhila - from that area the most numerous data, alt. 1420-1820 m (after 7 records), 7 stations. L 10: Nungwe trib., 1670 m, B&M 1079; O 4: Kalusembe R., 1270-1380 m, B&M 2838, Kapa Stm., 1720-1820 m, B&M 2802, Kipanda R„ 1550-1575 m, B&M 1198, 57 Libanda R., 1450 m, B&M 2919, Lukongolwe R., 1280-1360 m, B&M 2859, Mpa Stm., 1420 m, B&M 2947. Habitat. Terrestrial, in ravine and riparian forests in swampy places, near springs, on river and stream banks and on the scarps. Zambia - Kornas (1979): not noted. Athyrium schimperi Moug. ex Fee Distribution. Fig. 72. Very common, recorded from various parts of Upper Katanga - regions of Lualaba and Lufira Rivers, plateaux: Biano, Kundelungu and Muhila, regions of Luvua River, Lubumbashi and Mount Mupwelume (a part of those areas belongs to the “Katangan Copper Arc”), alt. 550-2000 m (after 66 records), 69 stations. G 10: near Kolwezi, 7 km from Luilu, 1250 m, B&M 348; G 11: ibid., Luilu Vai., 1160 m, B&M 343, Potopoto Stm., trib., Kolwezi - Musokatanda 30 km, 1425 m, B&M 336; H 10: Katentania R., 2 km from “Poste de Katentania”, 1500 m, B&M 707, 711, (Schelpe 1973, Kornas 1979); H 11: Kolwezi - Mamfwe 12 km, 1350 m, B&M 2030; I 10: Gule R„ 1450 m, B&M 538, Kalule Sud R„ 1500 m, B&M 612, 10 km S from Kansenia and from Kansenia Station, 1500 m, Lis. Mai. & Sym. 13496, (KORNAS 1979), Menge R., near Quaglia Farm, 1575 m, B&M 575, Mulamba R., fall, 1470 m, B&M 695; I 11: Fungurume “Hill V”, 1190 m, B&M 2058; I 12: Mindingi, Mai. 11764, Swambo, B&M 1142, Mai. 11773; J 11: Kanduba R., 6 km W from Kakontwe, 1330 m, B&M 825, Panda trib. near Likasi, 1150 m, Lis. 12151, (SCHELPE 1973, KORNAS 1979); J 12: Shinkolobwe, 1420 m, B&M 814; K 10: Kikuswa Stm., 6 km from Mwadingusha, B&M 399; K 12: Kamianga Stm., 5 km NW from Shilatembo, 1340 m, Mai. & Robbr. 1734, ibid., 5 km N from Shilatembo, 1200 m, B&M 290, Luafi Stm., Lubumbashi - Likasi 70 km, 1230 m, B&M 312, Milondo Stm., 3 km SW from Katanga, 1190 m, B&M 2016, Lukutwe Stm., 3 km ENE from Sofumwango Station, 1310 m B&M 850, Kabukende Stm., 2 km from the road to Sofumwango Station, 1300 m, B&M 862; K 13: Kafubu R., cascades, 1230 m, B&M 782, near Kafubu R„ Mukuen Hill, 1240 m, B&M 216, Kansaba, 1235 m, B&M 801, Kasombo Mine, 1300 m, B&M 66, Kibembe, 10 km N from Lubumbashi, 1300 m, B&M 208; L 8: Kimitondo Stm., Kibwe wa Sanga, 1730 m, B&M 1038; L10: Kapala R., 2 km N from Luishi, 1645 m, B&M 1010, Luanza R., 6 km N from Katshupa, 1670-1680 m, B&M 261, (SCHELPE 1973), Luanza R., spr. area, 1655 m, B&M 993; L 11: Kalembe R., 1550 m (Kornas 1979), Minwe Spr., 1620 m, B&M 963; M 4: Djania R„ 550 m, B&M 3121, Kalukenda Stm., 650 m, B&M 3110, Kamahasa Stm., 590 m, B&M 3167, Kasenga Stm., 590 m, B&M 3178, Kiminu R., 580 m, B&M 3189, Luvua R„ 590 m, B&M 3146, Musinoi R., 590 m, B&M 3198; M 5: Kamisamba R., 900 m, B&M 3308, Konkwa Stm., 690 m, B&M 3241, Lubalashi - Luvua confl., 680 m, B&M 3262, Lukongolwe Stm., 1000 m, B&M 3432, Mbula R., 700 m, B&M 3234, Muhezie R., 1000 m, B&M 3366, Mukalabafulue Stm., 900 m, B&M 3284, Mukungai R., 690 m, B&M 3229; N 5: Mukui R., 1300-1360 m, B&M 3580, Mukui trib., 1290-1360 m, B&M 3619; O 4: Kapondo R., 1500 m, B&M 1167, Kilumbi R., 1400 m, B&M 1855, Kyula Mt., 1670 m, B&M 751, Luilu trib., 1720-1820 m, B&M 1701, Mpa Stm., 1420 m, B&M 2937; P 5: Kasia R., Kieluzi - Mwela 58 40 km, 1780 m, B&M 1446, Lufuko R., 1790-1840 m, B&M 1483, Pepa, 2000 m (Kornas 1979); P 16: Akasansa R., spr. area, 1440 m, B&M 2556, Akapemfia R., spr. area, 1500 m, B&M 2617, Fibulukutu R., spr. area, 1450 m, B&M 2600, Kamakolongo R., 1440 m, B&M 2492, Kasamba R., 1250 m, B&M 2743, Kimbwi R„ spr. area, 1440 m, B&M 2579, Lungo R., 1370 m, B&M 2536, Lusemfwa R., 1500 m, B&M 2642, Tchitwichasonfu R., 1440 m, B&M 2454, Miloso R., Mupala, 1220 m, B&M 2752. Habitat. Terrestrial, mostly in riparian forests swampy in places, also in swampy, fringing and ravine dense forests, on river and seasonal stream banks and scarps, on forest edges, occasionally in disturbed forests, savannas, and on open sites, on rocks including mineralized outcrops and in “steppes” of copper mine region; found also on sandy soils. Zambia - Kornas (1979): 105. Ctenitis lanigera (Kuhn) Tardieu Distribution. Fig. 73. Very rare, Kalemie region near Lake Tanganyika, 1 station. O 1: Kabimba R., Lubandaye (Demaret 1955). Habitat. In riparian forest. Zambia - Kornas (1979): not noted. Diplazium zanzibaricum (Bak.) C. Chr. Distribution. Fig. 74. Very rare, Muhila Plateau, alt. 1390-1820 m, 2 (3) stations. O 4: Kalonga R„ 1390-1420 m, B&M 2879, Kapa Stm., 1720-1820 m, B&M 2795, Luilu trib. (maybe the same station as before), B&M 1704. Habitat. Terrestrial, in ravine forests, on ravine slopes and bottoms, near river rapids. Zambia - Kornas (1979): 105. Dryopteris athamantica (Kunze) Kuntze Distribution. Fig. 75. Very common, in various parts of Upper Katanga - regions of rivers Lualaba, Lufira and Luvua, plateaux: Biano, Kibara, Kunde-lungu, Marungu and Muhila, regions of Lubumbashi, Sakania, Mount Mupwe-lume and southern periphery of “Sakania Boot”, alt. 550-2400 (after 85 records), 92 stations. G 11: Kolwezi - Kapata 8 km, 1425 m, B&M 2037; H 10: Katentania R., 2 km from “Poste de Katentania”, 1500 m, B&M 706, env. Katentania (DE WlLDEMAN 1923, after SCHELPE 1973); I 9: Bisonso, 1400 m, B&M 449; I 10: Gule R„ 1450 m, B&M 541, 542, Kalule Sud R., 1500 m, B&M 614, E from Kalule Nord R., Shisinkwa, 1600 m (Schelpe 1973), Menge R., env. Quaglia Farm, 1575 m, B&M 559; I 11: Fungurume “Hill V”, 1190 m, B&M 2055; J 12: Shinkolobwe, 1420 m, B&M 808; K 7: Bana Ban-kasi Hill, Mitwaba - Manono 11 km, 1600 m, B&M 457; K 10: Kikuswa Stm., 6 km from Mwadingusha, B&M 404; K 12: Kamianga Stm., 5 km N from Shilatembo, 59 1200 m, B&M 292, Milondo Stm., 3 km SW from Katanga, 1190 m, B&M 2014; К 13: Fipango R., 1230 m, B&M 1117, Kafubu R., cascades, 1230 m, B&M 781, env. Kafubu R., Mukuen Mt., 1240 m, B&M 217, Kansaba Stm., Kaponda, 1235 m, B&M 802, Ka-ravia R., bridge, 1250 m, B&M 65, Kasapa, 1245 m, B&M 296, Kasombo Mine, 1300 m, B&M 69, Kasungami, 1225 m, B&M 174, ibid. Kamakoshi Stm., B&M 788, Kisanga, 1200 m (SCHELPE 1973), Lubumbashi - campus, 1250 m (SCHELPE 1973), 11 km from Lubumbashi, 1220 m (SCHELPE 1973, KORNAS 1979), 12 km NW from Lubumbashi, 1300 m (SCHELPE 1973), env. Nkingandu Mission, 1225 m, B&M 29; L 6: 10 km SSW from Kifinga (SCHELPE 1973); L 8: Kimitondo Stm., Kibwe wa Sanga, 1730 m, B&M 1033, Kwamba R., 1650 m, B&M 1043; L 10: Kapala R., 2 km N from Luishi, 1645 m, B&M 1021, 6 km N from Katshupa, 1680 m (SCHELPE 1973), Luanza Basin (MALAISSE 1975, KORNAS 1979), Luanza R., spr. area, 1655 m, B&M 1000; L 11: Minwe Spr., 1620 m, B&M 955; L 14: Kinsenda, 1330 m, B&M 2085; M 4: Djania R„ 550 m, B&M 3117, Kalukenda Stm., 650 m, B&M 3104, Kasenga Stm., 590 m, B&M 3172 b, Luvua R., 590 m, B&M 3140, Mangendwe Stm., 590 m, B&M 3179; M 5: Imakabanga Stm., 690 m, B&M 3211, Kakonkolwa Stm., 1190 m. B&M 3458, Kamian-ga R„ 720-800 m, B&M 3440, Kamimvu R., 690 m, B&M 3203, Kasa Mt., 650-690 m, B&M 3077, Konkwa Stm., 690 m, B&M 3238, Kyezimpala Stm., 690 m, B&M 3219, Luela R., B&M 3244, Lukongolwe Stm., 1000 m, B&M 3420, Mbopofu R., 1250 m, B&M 3338, Mbula, 700 m, B&M 3232, Muhezie R., 1000 m, B&M 3359, Mukungai R„ 690 m, B&M 3226; M 6: env. Kisele, B&M 83; N 4: Kajia R„ 1300-1350 m, B&M 3002; N 5: Kibondo R„ 1335 m, B&M 3026, Lualashi R„ 1300-1350 m, B&M 3050; N 15: Kapundu, 1230 m, B&M 1980, Lukangaba R„ 1230 m, B&M 1994, Munshi R„ 1225 m, B&M 1988; О 4: Kamuinga Stm., 1725 m, B&M 1657, Kapona Mt., 1750 m, B&M 1723, Katomia R., 1400 m, B&M 2955, Kilumbi R., Kilenge Rap., 1400 m, B&M 1849, Kisenga R„ 1425 m, B&M 1883, Laula R„ 1520 m, B&M 746, Libanda R„ 1450 m, B&M 2893, Mpa Stm., 1420 m, B&M 2928; О 15: Menda, 1230 m, B&M 1951; P 4: 2 km from Kitendwe, E from the road Moba - Pepa, 1930 m (SCHELPE 1973); P5: 12 km NNE from Kasiki, 2400 m (SCHELPE 1973), Misongo Mt., 2020 in, B&M 3866, Pepa, 2000 m (KORNAS 1979); P 16: Akangili R„ 1400 m, B&M 2440, Akapemfia R., spr. area, 1500 m, B&M 2621, Chinika R., spr. area, 1500 m, B&M 2662, Fibulukutu R., spr. area, 1450 m, B&M 2608, Kasala R., Mupala, 1245 m, B&M 2271, Kasamba R., 1250 m, B&M 2740, Kimbwi R., spr. area, 1440 m, B&M 2586, Lungo R., 1370 m, B&M 2533, Lusemfwa R., 1500 m, B&M 2641, 2648, Mabondo R., between Ifuna and Malashi, 1290 m, B&M 2329, Malashi R., 1275 m, B&M 2353, Miloso R„ Mupala, 1220 m, B&M 2753, Mubwe R„ 1350 m, B&M 2418, Nyamanda R„ 1290 m, B&M 2398, ibid. Spr., 1530 m, B&M 2679; 2 - precised location not available: Biano Plateau, 1550 m (KORNAS 1979), Kundelungu Plateau, 1650 m, B&M 229. Habitat. Terrestrial or lithophytic, mostly in riparian forests and at their edges, also in ravine and fringing forests, on the banks of rivers and also perennial and seasonal streams, on steep scarps; often in open woodlands (e.g. “miombo”) and savannas, on rocky slopes and summits (also on dolomitic rocks), in “steppe” vegetation (above 1900 m a.s.l.), near - or on - termite mounds, occasionally in swamps, sands and disturbed sites, such as gravel pits or road verges; in shade or in full sunlight. Zambia - Kornas (1979): 107. 60 Dryopteris inaequalis (Schlechtend.) Kuntze Taxonomic remark. Kornas (1979) stressed that this is “a highly critical complex, very much in need of taxonomical revision” and left specimens from Katanga (unlike that those from Zambia) without varietal distinction. For some voucher specimens (mentioned below separately) determination on species level was even difficult and is preliminary, in part because the existence of some new species is possible. It concerns mainly ferns from cupriferous soil. With reference to D. inaequalis Kornas distinguished (in card-index of Ka-tangan pteridophytes): “little divided form” (11 voucher specimens marked below by *) and “bipinnate form” (2 voucher specimens marked by **). Distribution. Fig. 76 (map includes typical and untypical forms). Very common, regions of Lualaba, upper Lufira and Luvua rivers, Manika Plateau and all high plateaux: Biano, Kibara, Kundelungu, Marungu and Muhila, regions of Lubumbashi and Mount Mupwelume, alt. 590-1880 m (after 52 records), 56 stations (68 stations together with specimens with uncertain determination). G 10: Luilu trib. near Kolwezi, 1130 m, B&M 356, Mupwiji R., 800 m, B&M 2175, Mushinji R., Kolwezi - Kyamasumba 70 km, 1010 m, B&M 2163; H 9: Kalule Sud R., Dikolongo Dam, 1400 m, B&M 9; I 9: Kalule Nord R., Dipera Falls, 1280 m, B&M 6; I 10: Kalule Sud R., 1500 m, B&M 616, 618, ibid. Biano Hotel, 1530 m (Schelpe 1973); Kipao R., 1590 m, B&M 645, 668, Pande R. Dam, Kansenia, 1400 m, B&M 136; I 11: Kimisombo Stm., Kakanda - Kambove 9 km, 1430 m, B&M 882, Luita, 1500 in, B&M 894', Tshilongo R., Kabwe, Mai. 11426', К 6: Mulumbi Esc., Mitwaba - Manono 52 km, 950 m, B&M 487', К 11: Kasala R., Mwadingusha - Koni 2 km, 1110 m, B&M 2130; К 12: Mwera R., 1150 m, B&M 1131, Sofumwango, 3 km ENE from the station, 1310 m, B&M 853; К 13: Kafubu R., Kasungami, 1210 m, B&M 293, Kibembe, 10 km N from Lubumbashi, 1300 m, B&M 207, Kiswishi Monastery, 1275 m, B&M 867; L 10: Luanza Basin, Katshupa (Malaisse 1975, Kornas 1979), Luanza R., 7 km NNE from Katshupa (SCHELPE 1973), Munyankara Stm., env. Gombela, 1060 m, B&M 388; L 11: Kaku R., 2 km N from Gombela, 1130 m, B&M 227, Lofoi R., between Touring Club and Kaloba Fall, 1080 m, B&M 1, Lutshipuka Falls, 1550 m, B&M 233, Masansa R., falls, 1500 m, B&M 266; M 4: Kamahasa Stm., 590 m, B&M 3162, Musinoi R., 590 m, B&M 3195; M 5: Musanikwa R„ 800-1000 m, B&M 3318; N 4: Kijiya R„ 1400-1425 m, B&M 1340*, Luhu R„ 1475-1500 m, B&M 1326; О 4: Kamabale Stm., 1350-1420 m, B&M 1633*, Kansimba R., 1425-1525 m, B&M 1151**, Kapa Stm., 1720-1820 m, B&M 2799*, Kapondo R., 1500 m, B&M 1179, Katwizia R„ 1400-1475 m, B&M 1187, Kisenga R., 1425 m, B&M 1875**, Kitampa Stm., spr. area, 1350-1400 m, B&M 1623*, Laula R„ 1240 m, B&M 719, Laula Rav., 1350-1400 m, B&M 1270*, Libanda R., 1450 m, B&M 2909, Luilu R., 1470 m, B&M 1794*, Mutambala Stm., 1500 m, B&M 1160, Nkala R., 1425 m, B&M 1347*; О 14: Luapula Vai., near Giraud Falls (KORNAS 1979); P 4: Kawa R., 1280-1450 m, B&M 3851, Kije R„ 750-1130 m, B&M 3839, Kisambila Stm., 750-1130 m, B&M 3824; P5: Kasia R„ Kieluzi - Mwela 40 km, 1780 m, B&M 1448, Kisanzo Stm., 1740 m, B&M 1428*, Lufuko trib., 7 km from Pepa, 1790-1840 m, B&M 1474, Lunanga R., 1640 m, B&M 1500*, Lunangwa - Kabwisi confl., 1820 m, B&M 1537*, Mufufu R., 1880 m, B&M 1555, Munamazi R., env. Kanipini, 1655 m, B&M 1518*; P 16: Bulanga R„ 1300 m, B&M 2366. 61 Habitat. Terrestrial, occasionally epiphytic, in narrow fringing forests, in riparian, swamp and dense forests in ravines and on escarpments, in open woodlands on rocky slopes, in wooded savannas - on river banks and scarps, rocks near waterfalls, at forest edges, in disturbed forest stands, also on cupriferous hills and in sandy soils, in shade and in open sites. Zambia - Kornas (1979): 108 - var. inaequalis. Dryopteris cfr. inaequalis Note. Specimens marked in KRA as “iuv. and dry”. Distribution. Found on Biano, Kibara and Muhila Plateaux and in the region of Luvua River, alt. 650-1850 m (after 7 records), 8 stations. I 11: 2-4 km S from Tshilongo Station, 1500 m, B&M 532; K 8: env. Lusinga, Upemba Natl. Park, 1510 m, B&M 81; M 4: Lusindoi R., 650 m, B&M 3091; O 4: Kilumbi R., Kilenge Rap., 1400 m, B&M 1832, Kipanda R., 1550-1575 m, B&M 1199, Musendemwa Stm., 1550-1850 m, B&M 1814; O 6: Kabinda Stm., B&M 1401; P 4: Mulobozi R., Kapupo Fall, 1020-1100 m, B&M 2764. Habitat. In riparian forests, ravines, and woodlands on the slopes, mostly on the rocks, found also on termite mound and - at 1 station - as epiphyte. Dryopteris sp. indet. - from D. inaequalis complex Distribution. Found near upper Lufira River, on Muhila Plateau and Mount Mupwelume, alt. 970-1900 m (after 4 records), 4 stations. K 11: Kalombe R., Mwashya - Koni 1 km, 970 m, B&M 2102; O 4: Kasenge Spr., Kibweza Mt., 1720-1900 m, B&M 1689, Kibola R„ 1400-1475 m, B&M 1219, 1220; P 16: Tchitwichasonfu R., 1440 m, B&M 2455. Habitat. Mostly in riparian forests and at river banks. Dryopteris kilemensis (Kuhn) Kuntze Distribution. Fig. 77. Very rare, Muhila Plateau and its western periphery, alt. 1300-1460 m (after 4 records), 4 stations. N 4: Kajia R„ 1300-1350 in, B&M 3006; N 5: Kibondo R„ 1335 m, B&M 3040; O 4: Lukonagulwa Stm., 1460 m, B&M 2821, Nkoko Stm., 1350 m, B&M 2987. Habitat. Terrestrial, in riparian forests, mostly near, or on river banks. Zambia - Kornas (1979): 109. Dryopteris sp. nova cuprophila ? - determination prelimirary Note. Suggestion of J. Kornas in KRA and in card-index of Katangan Pteri-dophytes. It concerns the mentioned below voucher specimen. Distribution and Habitat. Very rare, “Katangan Copper Arc”, 1 station. G 11: Dikuluwe, 1400 m, Mai. & Robbr. 2389, found on cupriferous hills. 62 Hypodematium crenatum (Forsk.) Kuhn Distribution. Fig. 78. Rare, Biano Plateau and “Katangan Copper Arc”, alt. 1130-1360 (after 5 records), 7 stations. I 10: Kalule Sud R. (SCHELPE 1973); I 11: Fungurume “Hill V”, 1130 m, B&M 2068, Luita, Mai. 10079, 11697, ibid, “cupriferous hill”, 1350 m, B&M 770; I 12: road Likasi - Kambove near bifurcation to Mindigi, 1320 m, B&M 771; J 11: 4-5 km N from Kakontwe, 1200 m, Mai. 10053, 10215, 11720, 6 km SSE from Kambove, 1360 m, B&M 836. Habitat. Lithophytic, in shallow ravine and cave, on calcareous rocks e.g. of Kakontwe, on siliceous rocks and hills with cupriferous formations. Zambia - Kornas (1979): 105. Polystichum fuscopaleaceum Alston (= P. seliferum (Forsk.) Moore var. fuscopaleaceum (Alston) Schelpe) Distribution. Fig. 79. Very rare, Marungu Plateau, 1 station. P5: Lunkole Stm., rav., Kieluzi - Mwela 45 km, 1890 m, B&M 1459. Habitat. Terrestrial, found in dense, in part disturbed forest of narrow ravine, on shady low situated site. Zambia - Kornas (1979): not noted. Tectaria gemmifera (Fee) Alston Distribution. Fig. 80. Quite common, mostly on the plateaux: Biano, Kun-delungu, Muhila (most numerous data) and Marungu, alt. 1080-1890 (after 11 records), 12 stations. 19: Kayo Falls, 1250-1310 m, B&M 412; I 10: Pande R., fall, 1300 m, B&M 590; L 10: Lofoi Vai., Kaloba Fall, 1200 m, B&M 168; L 11: ibid, between Touring Club and Kaloba Fall, 1080 m, B&M 929; N 1: Tendele R., Tugulu (Demaret 1955); O 1: Lu-bandaye R., Kabimba, 1200 m (Demaret 1955); O 4: Kapa Stm., 1720-1820 m, B&M 2803, Laula R„ rav., 1250 m, B&M 725, ibid. 1350-1400 m, B&M 1266,1274, Libanda R., 1450 m, B&M 2908, Luilu trib., 1720-1820 m, B&M 1708; P 5: Lunkole Stm., Kieluzi - Mwela 45 km, 1890 m, B&M 1463. Habitat. Terrestrial or lithophytic, in riparian and ravine dense forests, on scarps and river banks, on rocks situated in rivers and near waterfalls. Zambia - Kornas (1979): 106. Tectaria sp. indet. Note. According to the remarks of A. Bodenghien and F. Malaisse in KRA specimens “non gemmiferes”. Distribution. Found in the region of Lubule River and of Lake Moero, alt. 970-1240 m, 3 stations. 63 M 7: Kamitaba R., 990-1240 m, B&M 4475, Mukolowanga R„ 990-1100 m, B&M 4496; N 7: Mibiluko R., 970-1050 m, B&M 4554. Habitat. Ravine dense forests, on rocky scarps. Gleicheniaceae Dicranopteris linearis (Burm. f.) Underw. Note. Voucher specimen: Lisowski 10100 is identified as D. linearis var. linearis. Distribution. Fig. 81. Common, region of rivers Lualaba and Luvua, plateaux: Biano, Kibara, Kundelungu, Muhila and Marungu, regions of Lubumba-shi and of Mount Mupwelume, alt. 810-1900 m (after 22 records), 25 stations. G 10: Mutendele R., 810 m, B&M 367; G 11: Nzilo Rav., N from Kolwezi, 1180 m, Lis. 10100 - var. linearis (KORNAS 1979); H 10: Katentania, 1560 m (SCHELPE 1973), 2 km from “poste de Katentania”, 1500 m, B&M 703; 110: Kisamoya R., 1500 m, B&M 633; K 6: Bana Bankasi Hill, Mitwaba - Manono 11 km, 1600 m, B&M 455, Mitwaba - Sermikat 11 km, 1600 m (SCHELPE 1973); K 12: Luafi Stm., Lubumbashi - Likasi 70 km, B&M 319; K 13: 15 km SE from Lubumbashi, 1150 m (SCHELPE 1973); L 10: 6 km N from Katshupa (SCHELPE 1973), Luanza Basin (MALAISSE 1975, KORNAS 1979); L 11: Lutshipuka Falls, 1520 m, B&M 243; L 13: Munama R., Kasamba, 1220 m. B&M 1583; N 5: Lualashi R„ 1300-1350 m, B&M 3053, Mukui trib., 1290-1360 m, B&M 3611, 3641; O 4: Kamuinga R., 1725 m, B&M 1654, Kansimba trib., 1490 m, B&M 1763, Katomia R„ 1400 m, B&M 2959, Libanda R„ 1450 m, B&M 2897; P 4: Mulobozi R., Kapupo Fall, 1020-1100 m, B&M 2763; P 5: near Mufufu R., 1900 m, B&M 1573; P 16: Akeshimbi R., 1300 m, B&M 2373, Bulanga R., 1275 m, B&M 2355, Fibulukutu R., spr. area, 1450 m, B&M 2592, Malashi R., 1275 m, B&M 2340. Habitat. Terrestrial, in riparian and ravine dense forests and at their edges, on slopes with seeping water, on river banks, sandy and rocky scarps and sands, in wooded savannas on rocky slopes or on alluvial sites, on wet places between rocks and boulders, found also along the roads and in the thickets on artificial ditch; in shade and in sunny places. Zambia - Kornas (1979): 74 (only D. linearis var. linearis). Hymenophyllaceae Note. The studies about Hymenophyllaceae in Central Africa (including Upper Katanga), with maps of species distribution, are published in two separate papers by Kornas (1993 b, 1994). Hymenophyllum hirsutum (L.) Swartz (= H. ciliatum Sw. (pro parte)) Distribution. Fig. 82. Quite common, mostly on the plateaux: Biano, Kibara, Kundelungu and Muhila, also in the regions of Luvua River and Plain of 64 Lubumbashi, alt. 1200-1680 m (after 13 records), 13 stations; nearly all of them are quoted by Kornas (1993 b). 110: Kalule Sud R., Biano Hotel, 1560 m (Schelpe 1973), Kipao R„ 1590 m, B&M 654, (KORNAS 1993 b), Menge R., near Quaglia Farm, 1575 m, B&M 580, (KORNAS 1993 b); K 6: Kanda Stm., Mitwaba - Manono 34 km, 1400 m, B&M 461, (KORNAS 1993 b); K 12: Luafi Stm., rav., near Mangombo, Lubumbashi - Likasi 70 km, 1200-1300 m, B&M 316, 2088, Mai. & Robbr. 2129, (KORNAS 1993 b), Lubumbashi - Likasi 97 km, 1200 m, Mai. 10058, (KORNAS 1993 b); L 10: Luanza R., spr. area, 1655 m, B&M 994, (KORNAS 1993 b), Luanza R., 6 km N from Katshupa, 1660-1680 m, B&M 264, (MALAISSE 1969 as H. ciliatum, Schelpe 1973, KORNAS 1993 b), ibid., 7 km NNE from Katshupa, 1560 m (MALAISSE 1969 as H. ciliatum, SCHELPE 1973); N 4: Mukui trib., 1300-1360 m, B&M 3710, (Kornas 1993 b as Kasenga R.); N 5: Kasenga R., 1300-1340 m, B&M 3562, Mukui R. and trib., 1285-1360 m, B&M 3600, 3685, (Kornas 1993 b, two last numbers as Kasenga R.); O 4: Kisenga R., 1425 m, B&M 1862, (Kornas 1993 b). Habitat. Epiphytic or litophytic, in riparian and fringing forests, in ravine dense forests, on the slopes, wet (occasionally sandy) scarps, at river banks on the rocks and moistened sand with gravel. Zambia - Kornas (1979): not noted. For distribution in Central Africa see Kornas (1993 b). Hymenophyllum kuhnii C. Chr. (= H. polyanthos Sw. var. kuhnii (C. Chr.) Schelpe) Distribution. Fig. 83. Very rare, Kibara Plateau, Upemba National Park, 1 station. K 7: Katongo, 1750 m, de Witte 3720, (Kornas 1993 b). Habitat. Small epiphyte in riparian forest. Zambia - Kornas (1979): species not noted in that country but collected on the Nyika Plateau on Malawi side (Kornas I. c.: 72 - after Schelpe 1970 - H. polyanthos var. kuhnii). For distribution in Central Africa see Kornas (1993 b). Trichomanes chevalieri Christ Note. All stations are quoted after Kornas (1994), according the text and map in this publication (citations ommited below) and after some additional information from unpublished card-index. Data of Kornas are based on Brussels Herbarium (BR). Distribution. Fig. 84. Very rare in Upper Katanga, found near Lake Tanganyika, 1 station. O 1: Tendele R., Tugulu, in riparian forest, on moist soil. Beyond Upper Katanga, nearby its border 11 stations, mostly in the region of River Lulua and slightly to the East - B 8: Kalongani Stm., env. Kalozia, 960 m, Kamuno Stm., trib. of Lumbadi, 970 m, Kazaza R., trib. of Mulendo, Muteba, 980 m, Luandu R., trib. of Lulua, Mutondokania, 960 m, Lufwazeu R., trib. of Lulua, env. Shimbu, 970 m, 65 Lumbadi R., 970 m, Nakabam R., trib. of Kazaza, Muteba, 1000 m, Ringambo Stm., 960 m; E 5: Kaniama - Haut-Lomami, 900 m; G 7: Kamina R., 3 km S from Kamina; J 1: Katundu, 45 km SE from Kongolo. Habitat. Mostly epiphytic on tree trunks, in moist environment of riparian forests. Zambia - Kornas (1979): not noted. Trichomanes clarenceanum Ballard Distribution. Fig. 85. Very rare, region of Lualaba River, 1 station. G 10: Mutendele R., Kolwezi - Luena 114 km, 785 m, B&M 2039, (KORNAS 1994). Habitat. Riparian forest, on the foots of tree trunks on steep slope near the water. Zambia - Kornas (1979): not noted. For distribution in Central Africa see Kornas (1994). Trichomanes cfr.fallax Christ Note. J. Kornas called the attention in KRA on tripinnate leaves whereas in typical T. fallax the leaves are only bipinnate - therefore the determination neads revision. Distribution. Fig. 86. Very rare, Biano Plateau, 1 station. I 10: Kadiva Falls, 1200 m, B&M 604. Habitat. Found under some shelter in the waterfall. Zambia - Kornas (1979): not noted. Trichomanes frappieri Cordem. Distribution. Fig. 87. Very rare, Muhila Plateau, 1 station. O 4: Kilumbi R., Kilenge Rap., 1450-1500 m, B&M 1310 a, (KORNAS 1994). Habitat. Lithophytic and epiphytic in riparian forest. Zambia - Kornas (1979): not noted. For distribution in Central Africa see Kornas (1994). Trichomanes giganteum Bory Distribution. Fig. 88. Very rare, Biano Plateau, 1 station. I 11: Kansenia, Kadiva Falls, 1400 m (SCHELPE 1973). Habitat. Terrestrial or epiphytic - according Johns (1991). Found in, and below the waterfall, under overhanging rocks and in rock crevices. Zambia - Kornas (1979): not noted. Trichomanes inopinatum (Pic. Ser.) J. E. Burrows Distribution. Fig. 89. Rare, Biano, Muhila and Marungu Plateaux, alt. 1220-1655 m (after 3 records), 6 stations. All stations mentioned below are quoted by Kornas (1994), some of them are not represented by specimens in KRA. 66 I 9: Dipera Falls, 1220 m, B&M 426, (KORNAS 1994), Kayo Falls, env. Lubudi (Kornas 1994); O 4: Kabwa R. (KORNAS 1994), Laula R„ rav. 1350-1400 m, B&M 786, 1264, 1280, (KORNAS 1994); P 5: Munamazi R., env. Kanipini, 1655 m, B&M 1530, (Kornas 1994). Habitat. Epiphytic or lithophytic, in ravine dense forest, in fringing forest between steep slopes, on steep rocky scarp near waterfall, on the basis of tree trunks, and occasionally on the dry rocks. Zambia - Kornas (1979): not noted. For distribution in Central Africa see Kornas (1994). Trichomanes melanotrichum Schlechtend. (= T. pyxidiferum L. var. melanotrichum (Schlechtend.) Schelpe) Distribution. Fig. 90. Very common, region of Lualaba River, plateaux: Biano, Kundelungu, Marungu and Muhila - with its periphery (most numerous sites), region of Luvua River, alt. 680-1850 m (after 34 records), 39 stations; 5 other station beyond Upper Katanga. All those stations are mentioned in publication of Kornas (1994), therefore this quotation is ommited below with exception of stations without other information. G 10: Mushinji R., Kolwezi - Kyamasumba 70 km, 1010 m, B&M 2164-, I 10: Kipao R., 1590 m, B&M 655, 666, Menge R., env. Quaglia Farm, 1575 m, B&M 581, Mulamba R., 1470 m, B&M 686; I 11: Kolwezi - Kamina road, Schm. 1778, Tshilongo Stm., Mukabe, 1100 m, Mai. 10120, Tshilongo Stm., 1300 m, B&M 330-, L 9: Vibyo R., 1670 m, B&M 1065; L 11: Katwe R„ 1645 m, B&M 1097, Minwe Spr., 1620 m, B&M 964; M 5: Kamianga - Lubalashi confl., 720-800 m, B&M 3457, Kamisamba R., 900 m, B&M 3315, Kamabale Stm., 1200 m, B&M 3489, Lubalashi - Luvua confl., 680 m, B&M 3271, Mukalabafulue Stm., 900 m, B&M 3292; N 4: Kajia R., 1300-1350 m, B&M 3023, LuhuR., 1475-1500 m, B&M 1332; N 5: Kibondo R„ 1335 m, B&M 3048, Lualashi R., 1300-1350 m, B&M 3074, Mukui R., 1300-1360 in. B&M 3603, Mukui trib., 1290-1360 m, B&M 3637, 3666; O 4: Kalonga R„ 1390-1420 m, B&M 2887, Kansimba, Lis. 12101, Lumbadi R. (KORNAS 1994), Kapa Stm. B&M 2810, Kapondo R., 1500 m, B&M 1176, Katomia R., 1400 m, B&M 2978, Kibola R„ 1400-1475 m, B&M 1214, Kilumbi R„ Kilenge Rap., 1450-1500 in, B&M 1310. Kisenga R., 1425 m, B&M 1879, Laula Rav., 1350-1400 in, B&M 1248, Libanda R„ 1450 m. B&M 2927, Lukonagulwa Stm., 1460 in, B&M 2828, Lukongolwe R., 1280-1360 m, B&M 2869, Musendemwa Stm., spr. area, 1550-1850 m, B&M 1829, Nkala R., 1425 in, B&M 1353, Nkoko Stm., 1350 m, B&M 3000; O 4?: Muhila Plateau (KORNAS 1994); P5: Mufufu R„ 1880 m, B&M 1563. Beyond Upper Katanga, near of its border, 5 stations in the region of Lulua River and of Kaniama (after KORNAS 1994, supplemented by unpublished card index, data from BR) - B 8: Luandu R., trib. of Lulua, Mutondonkania 960 in, Lumbadi R., 970 in, Mulendo R. trib. of Lulua, 920 m, Ringambo Stm., 960 m; E 5: Kaniama - Haut-Lomami. Habitat. Epiphytic or occasionally lithophytic. Mostly in riparian forests (in part narrow, closed by slopes) and in ravine dense forests, on the rocks and rocky scarps, on river banks and in swamp forests near sources. Zambia - Kornas (1979): 72 (T. pyxidiferum var. melanotrichum). For distribution in Central Africa see Kornas (1994). 67 Trichomanes radicans Sw. Distribution. Fig. 91. Very rare, Biano Plateau, I station. I 10: Kansenia, Kadiva Falls (KORNAS 1994). Habitat. Usually low-level epiphyte in moist forests. Zambia - Kornas (1979): not noted. For distribution in Central Africa see Kornas (1994). Trichomanes rigidum Sw. (= T. cupressoides Desv.) Distribution. Fig. 92. Quite common, mainly Biano and Kundelungu Plateaux, also Kibara Plateau and regions of Luvua River and of Lake Tanganyika, alt. 1530-1700 m (after 10 records), 12 stations; they are - with two exceptions - quoted by Kornas (1994). I 10: Kipao R., 1530-1630 m, B&M 646, 653, 661, (KORNAS 1994), Menge R., near Quaglia Farm, 1575 m, B&M 579, (Kornas 1994), Mulamba Road, 4 km from Mulam-ba Falls, alt. 1700 m, B&M 145, (KORNAS 1994); J 7: Upemba Natl. Park, Buye-Bala R., 1750 m, de Witte 3633, (KORNAS 1994); K 7: Upemba Natl. Park, Kalumengongo R., 1830 m, de Witte 5958, (KORNAS 1994); L 10: Kapala R., 2 km N from Luishi, 1645 m, B&M 1024, (Kornas 1994), Luanza Basin (Malaisse 1975, Kornas 1979), Luanza R., spr. area, Katshupa, 1675 m, B&M 991, (KORNAS 1994), Luanza Vai., 6 km N from Katshupa, 1680-1690 m, B&M 255, 265 a, (SCHELPE 1973, KORNAS 1994), Nungwe trib., 1670 m, B&M 1075, (Kornas 1994); N 6: 33 km ESE from Kasama (SCHELPE 1973); O 1: Lubandaye R., Kabimba, 1200 m (DEMARET 1955). Habitat. Terrestrial, occasionally epiphytic (also on the above-ground tree roots), usually in riparian, fringing and ravine dense forests, on steep river banks and scarps, also in swamp forests near springs; in shade, occasionally in the open, e.g. in disturbed forest stands. Zambia - Kornas (1979): 73. For distribution in Central Africa see Kornas (1994). Trichomanes sp. indet. Distribution. Specimens found on Biano and Muhila Plateaux, at 2 stations. I 9: Kayo Falls, 1250-1310 m, B&M 411; O 4: Kabwa R„ 1450 m, B&M 1229. Habitat. Epiphytic or lithophytic, on decayed trunk and on rocky scarp in riparian forest. Lomariopsidaceae Bolbitis acrostichoides (Afz. ex Sw.) Ching Distribution. Fig. 93. Very rare, region of Lualaba River and Muhila Plateau, alt. 785-1400 m (after 2 records), 2 stations. 68 G 10: Mutendele R., Kolwezi - Luena 114 km, 785 m, B&M 2043', О 4: Laula R., 1350-1400 m, B&M ¡273. Habitat. Terrestrial, in riparian and ravine dense forests, situated in part on steep slopes - there on rocks in places with seeping water. Zambia - Kornas (1979): not noted. Bolbitis auriculata (Lam.) Alston Distribution. Fig. 94. Very rare, near Lake Tanganyika, 1 station. О 1: Lubandaye R., Kabimba, 1200 m (Demaret 1955). Habitat. Terrestrial, in riparian forest on wet sites. Zambia - Kornas (1979): not noted. Bolbitis heudelotii (Bory ex Fee) Alston Distribution. Fig. 95. Common, region of Lualaba River, plateaux: Biano and Muhila with its western periphery, region of Luvua River and of lakes Tanganyika and Моего, Plain of Lubumbashi, alt. 720-1820 m (after 23 records), 23 stations. G 10: Mutendele R., 810 m, B&M 371, ibid. Kolwezi - Luena 114 km, 785 m, B&M 2041', I 10: Kadiva Falls, upstream of Kansenia, 1300 m, B&M 608; К 13: Kibembe R., Lukuni, 1250 m (SCHELPE 1973); M 5: Kamianga - Lubalashi confl., 720-800 m, B&M 3455; N 4: Kajia R„ 1300-1350 m, B&M 3021, Kitebe R„ 1300-1350 m, B&M 3813, Luiboso R., 1300 m, B&M 3787, Nungu R„ 1300 m, B&M 3763; N 5: Kibondo R„ 1335 m, B&M 3049, Mukui R., 1300-1360 m, B&M 3601, Mukui trib., 1285-1360 m, B&M 3660, 3682, 3706; N 7: Kasanza R„ 950-1050 m, B&M 4636, Mbikwa R„ 975-1050 m, B&M 4581, Mulonde R., near Kanfwa, 1030 m, B&M 1372, Mutemuna R., Kyankalamu - Pweto 77 km, 1290 m, B&M 1395; О 4: Kamabale Stm., 1350-1420 in, B&M 1636, Kapa Stm., 1720-1820 m, B&M 2807, Kapondo R„ 1500 m, B&M //66, 1172, Kisenga R„ 1425 m, B&M 1877, 1882, Laula Fall, 1400-1450 m, B&M 1281, Laula Stm., rav., 1450-1550 m, B&M 1283, 1288; Q 6: Mukonge R., env. Sunkutu, 1100 m, B&M 4038; R 6: Kalembwe R„ 730-920 m, B&M 4025. Habitat. Amphiphytic, on the rocks in rivers and waterfalls, mostly amongst riparian forests, also in ravines. Zambia - Kornas (1979): 109. Elaphoglossum acrostichoides (Hook, et Grev.) Schelpe Distribution. Fig. 96. Very rare, Kundelungu Plateau, 1 station. L 10: Luanza R., 5 km from spring and 6 km NNE from Katshupa, 1580 m (Schelpe 1973). Habitat. Epiphyte, in narrow fringing forest. Zambia - Kornas (1979): 110 “recorded from Zambia by Schelpe (1969) but not confirmed later”. 69 Elaphoglossum angulatum (Bl.) Moore Distribution. Fig. 97. Very rare, region of Mount Mumpwelume, 1 station. P 16: Kasamba R., 1250 m, B&M 2742. Habitat. Found in shallow ravine, 3 m deep. Zambia - Kornas (1979): not noted. Elaphoglossum chevalieri Christ Distribution. Fig. 98. Rare, region of upper Lufira River and of Lubumba-shi, 1150-1330 m (after 5 records), 5 stations. J 11: 4 km N from Kakontwe, 1200 m, Mai. 10055, Kanduba R., 8 km W from Kakontwe, 1330 m, B&M 830; K 12: Luafi Stm., Likasi - Lubumbashi 70 km, 1200-1230 m, B&M 2089, 314, Lubumbashi - Likasi 68 km, 1150 m, Mai. 10210, 3 km ENE from Sofumwango Station, 1310 m, B&M 844. Habitat. Terrestrial, in ravines, mostly with dense forests and seasonal streams, on steep scarps; found also on calcareous outcrops (near Kakontwe). Zambia - Kornas (1979): not noted. Elaphoglossum hybridum (Bory) Brack Distribution. Fig. 99. Very rare, Muhila Plateau, 1 station. O 4: Kilumbi R., Kilenge Rap., 1450-500 m, B&M 1313, 1848. Habitat. Lithophytic, found in riparian forest, on rocky scarp and banks of the river. Zambia - Kornas (1979): not noted. Elaphoglossum petiolatum (Sw.) Urban subsp. salicifolium (Willd. ex Kaulf.) Schelpe (= E. salicifolium (Willd. ex Kaulf.) Alston) Distribution. Fig. 100. Very common, regions of rivers Lufira, Luvua and upper Luapula, plateaux: Biano, Kibara, Kundelungu, Muhila and Marungu, Plain of Lubumbashi and south-eastern periphery of “Sakania Boot” with Mount Mupwelume, alt. 720-1840 m (after 60 records), 62 stations. 110: Gule R., 1450 m, B&M 537, Menge R„ env. Quaglia Farm, 1575 m, B&M 568, Mulamba Falls, 1450 m, B&M 677; J 11: Kanduba R., 8 km W from Kakontwe, 1330 m, B&M 829; K 6: Kyowe R„ Mitwaba - Manono 48 km, 1140 m, B&M 488; K 10: Kiku-swa Stm., 6 km from Mwadingusha, 1325 m, B&M 401; K 12: Kabukende Stm., 2 km from the road to Sofumwango Station, 1300 m, B&M 859, Kamianga Stm., 5 km N from Shilatembo, 1500 m, B&M 283, Lukutwe Stm., 3 km ENE from Sofumwango Station, 1310 m, B&M 842, Milondo Stm., 3 km SW from Katanga, 1190 m, B&M 2011; L 9-10 ?: Kalunda R., 1590 m (KORNAS 1979); L 10: Kapala R., 2 km N from Luishi, 1645 m, B&M 1026, Luanza Basin (Malaisse 1975, Kornas 1979); L 11: Kalembe R„ 1550 m (KORNAS 1979), Lutshipuka Falls, 1500-1520 m, B&M 235, 241 70 bis, 244, 11387, Masansa Fall, 1500 m, B&M 273; M 5: Kamabale Stm., 1200 m, B&M 3482, Kimatambo Stm., 1350 m, B&M 3389, Lukongolwe Stm., 1000 m, B&M 3427, Muhezie R., 1000 m, B&M 3363, 3373, Tumengele R., 900 m, B&M 3407, Kamianga - Lubalashi confl., 720-800 m, B&M 3447; M 10: Luanza R., 15 km WSW from Kabiashia, 1420 m (SCHELPE 1973); N 4: Kitebe R., 1300-1350 m, B&M 3801, Luiboso R., 1300 m, B&M 3785, Nungu R., 1300 m, B&M 3757; N 5: Kabanga R., 1175— 1200 m, B&M 3519, Kaha R„ 1290-1320 m, B&M 3530, Kanianga trib., 1100-1150 m, B&M 3506, Kasenga R„ 1300-1340 m, B&M 3559, Kibondo R„ 1335 m, B&M 3038, Kilungu Stm., 1075-1100 m, B&M 3511, Lualashi R., 1300-1350 m, B&M 3067, ibid, near spr., 1350-1410 m, B&M 3544, Mukui R., 1300-1360 m, B&M 3577, Mukui trib., 1285-1360 m, B&M 3614, 3646, 3675, 3697; O 1: Lubandaye R„ Kabimba, 1200 m (Demaret 1955); O 2: Koki R„ Mulange (Demaret 1955); O 4: Kalonga R., 1390— 1420 m, B&M 2876, Kalusembe R., 1270-1380 m, B&M 2839, Katomia R., 1400 m, B&M 2975, Kilumbi R„ Kilenge Rap., 1400 m, B&M 1846, ibid. 1450-1500 m, B&M 1312, Kisenga R., 1425 m, B&M 1861, Libanda R„ 1450 m, B&M 2911, Nkoko Stm., 1350 m, B&M 2994; O 15: Kiangulube Stm., 1300m,B